Endoceratoidea
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Endoceratoidea
Endocerida is an extinct nautiloid order, a group of cephalopods from the Lower Paleozoic with cone-like deposits in their siphuncle. Endocerida was a diverse group of cephalopods that lived from the Early Ordovician possibly to the Late Silurian. Their shells were variable in form. Some were straight (orthoconic), others curved (cyrtoconic); some were long (longiconic), others short (breviconic). Some long-shelled forms like ''Endoceras'' attained shell lengths close to . The related ''Cameroceras'' is anecdotally reported to have reached lengths approaching , but these claims are problematic. The overwhelming majority of endocerids and nautiloids in general are much smaller, usually less than a meter long when fully grown. Morphology Endocerids had a relatively small body chamber as well as a proportionally large siphuncle, which in some genera reached nearly half the shell diameter. This suggests that much of the visceral mass may have been housed within the siphuncle itself ...
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Nautiloidea
Nautiloids are a group of marine cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living ''Nautilus'' and ''Allonautilus''. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms ( orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day. In a broad sense, "nautiloid" refers to a major cephalopod subclass or collection of subclasses (Nautiloidea ''sensu lato''). Nautiloids are typically considered one of three main groups of cephalopods, along with the extinct ammonoids (ammonites) and living coleoids (such as squid, octopus, and kin). While ammonoids and coleoids are monophyletic clades with exclusive ancestor-descendant rela ...
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Cephalopod
A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by ''Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been ident ...
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Cameroceras
''Cameroceras'' ("chambered horn") is a genus of extinct, giant orthoconic cephalopod that lived mainly during the Ordovician period. It first appears during the middle Ordovician, around 470 million years ago, and was a fairly common component of the fauna in some places during the period, inhabiting the shallow seas of Laurentia, Baltica and Siberia.Frey, R.C. 1995. U.S. Geological Survey, p.73 Its diversity and abundance became severely reduced following the Ordovician–Silurian extinction events, and the last remnants of the genus went extinct sometime during the Wenlock. In older literature ''Cameroceras'' was treated as the largest nautiloid with shell length about or even , however the large-sized species is no longer belong to genus ''Cameroceras'' but ''Endoceras giganteum'', and the -long specimen is highly doubtful. Description ''Cameroceras'' is a cephalopod, a taxon of molluscs that includes the octopuses, squids and cuttlefish. From comparison with living cephalo ...
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Bisonocerida
Bisonocerida is an order of Ordovician to Silurian nautiloid cephalopods. Members of this order were originally placed in the order Endocerida, but later investigation argued that this broad usage of Endocerida was a polyphyletic assemblage encompassing two different groups of independent origin. Bisonocerida was differentiated from Endocerida in 2012 in order to resolve this issue. Bisonocerids are similar to endocerids in many respects. The siphuncle was broad and positioned ventrally in the shell, which ranged in shape from cyrtoconic brevicones (curved and short) to rare orthoconic longicones (straight and long). The inner surface of the siphuncle contains endosiphuncular deposits, which help distinguish the two orders. In both bisonocerids and endocerids, the endosiphuncular deposits are conical in shape ("endocones"), concreted from the rim of the siphuncle and tapering towards the apex of the shell. Both orders possess simple perforate endocones, with a hole at the tip of ...
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Orthoceratoidea
Orthoceratoidea is a major subclass of nautiloid cephalopods. Members of this subclass usually have orthoconic (straight) to slightly cyrtoconic (curved) shells, and central to subcentral siphuncles which may bear internal deposits. Orthoceratoids are also characterized by dorsomyarian muscle scars (a small number of large scars concentrated at the top of the body chamber), extensive cameral deposits, and calciosiphonate connecting rings with a porous and calcitic inner layer. Currently, Orthoceratoidea comprises the orders Riocerida, Dissidocerida, Actinocerida, Pseudorthocerida, Lituitida and Orthocerida. Orthocerida is a noteworthy paraphyletic order which is ancestral to the major cephalopod groups such as the extinct ammonoids and living coleoids (cephalopods without external shells, including squids, octopus, cuttlefish, etc.). Taxonomy As a superorder, Orthoceratiodea was one of six superorders within the Nautiloidea, the others being the Plectronoceratoidea (= Ellesm ...
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Endoceras
''Endoceras'' (Ancient Greek for "inner horn") is an extinct genus of large, straight shelled cephalopods from the Middle and Upper Ordovician that gives its name to the Nautiloid order Endocerida. The cross section in the mature portion is slightly wider than high, but is narrower laterally in the young. Sutures are straight and transverse. ''Endoceras'' has a large siphuncle, located close to the ventral margin, composed of concave segments, especially in the young but which may be tubular in the adult stage. Endocones are simple, subcircular in cross section, and penetrated by a narrow tube which may contain diaphragms reminiscent of the Ellesmerocerid ancestor. ''Endoceras'' was named by Hall in 1847. Distribution is widespread, especially in North America and Europe; and fossils have been found in Australia. ''Endoceras'' is similar to ''Cameroceras'', the two may be synonymous, but differs from the genus '' Nanno'' in that the siphuncle in ''Nanno'' fills the entire api ...
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Endoceras Sp
''Endoceras'' (Ancient Greek for "inner horn") is an extinct genus of large, straight shelled cephalopods from the Middle and Upper Ordovician that gives its name to the Nautiloid order Endocerida. The cross section in the mature portion is slightly wider than high, but is narrower laterally in the young. Sutures are straight and transverse. ''Endoceras'' has a large siphuncle, located close to the ventral margin, composed of concave segments, especially in the young but which may be tubular in the adult stage. Endocones are simple, subcircular in cross section, and penetrated by a narrow tube which may contain diaphragms reminiscent of the Ellesmerocerid ancestor. ''Endoceras'' was named by Hall in 1847. Distribution is widespread, especially in North America and Europe; and fossils have been found in Australia. ''Endoceras'' is similar to ''Cameroceras'', the two may be synonymous, but differs from the genus '' Nanno'' in that the siphuncle in ''Nanno'' fills the entire api ...
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Endoceratidae
Endoceratidae is a family of large to very large straight shelled nautiloid cephalopods belonging to the order Endocerida that lived during the Middle and Late Ordovician. They include the largest known Paleozoic invertebrates, represented by '' Endoceras'' and ''Cameroceras''.Teichert C, (1964). Endoceratoidea; Treatise on Invertebrate Paleontology, ''Part K, Mollusca 3''.Flower R. H. (May 1958). "Some Chazyan and Mohawkian Endoceratida", ''Jour Paleo'' 32 (2) pp 433-458 Description Endocerids are characterized by large, generally ventral siphuncles with simple endocones which lack the more complex structures associated with the Proterocameroceratidae and their derivatives. Endocones, nested cone-shaped deposits characteristic of the Endocerida, form in the apical portion of the siphuncle where they balanced the weight of the animal in the body chamber at the opposite end. This helped assure a horizontal orientation while the endoceratid animal was alive. The septal necks are ...
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Siphuncle
The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and ''Spirula''. In the case of the cuttlefish, the siphuncle is indistinct and connects all the small chambers of that animal's highly modified shell; in the other cephalopods it is thread-like and passes through small openings in the septa (walls) dividing the camerae (chambers). Some older studies have used the term siphon for the siphuncle, though this naming convention is uncommon in modern studies to prevent confusion with a mollusc organ of the same name. Function The siphuncle is used primarily in emptying water from new chambers as the shell grows. To perform this task, the cephalopod increases the saltiness of the blood in the siphuncle, and the water moves from the more dilute chamber into the blood through osmosis. At the sam ...
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Nekton
Nekton or necton (from the ) refers to the actively swimming aquatic organisms in a body of water. The term was proposed by German biologist Ernst Haeckel to differentiate between the active swimmers in a body of water, and the passive organisms that were carried along by the current, the plankton. As a guideline, nektonic organisms have a high Reynolds number (greater than 1000) and planktonic organisms a low one (less than 10). However, some organisms can begin life as plankton and transition to nekton later on in life, sometimes making distinction difficult when attempting to classify certain plankton-to-nekton species as one or the other. For this reason, some biologists choose not to use this term. History The term was first proposed and used by the German biologist Ernst Haeckel in 1891 in his article ''Plankton-Studien'' where he contrasted it with plankton, the aggregate of passively floating, drifting, or somewhat motile organisms present in a body of water, primarily t ...
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Allotrioceras
''Allotrioceras'' is a tubular fossil from the Middle Ordovician of the state of New York, collected by Rousseau H. Flower; included by him in the Endocerida and placed in a new family, the Allotrioceratidae. ''Allotrioceras'' is characterized by a lateral pair of subequal conical structures, resembling the endocones of endocerids, separated by a straight partition that extends more than half way across from either the dorsal or ventral side, as perceived, and runs along the length. What remains is thought to represent a siphuncle, which ranges from about 8 millimeters (0.3 in) to about 15 millimeters (0.6 in) in diameter. On the basis of diagnosis, inclusion of ''Allotrioceras'' and the Allotrioceratidae, for which it is the type, in the endocerida seems at best tentative. Some endocerids, derived from the Proterocameroceratidae, ''Najaceras'', '' Cacheoceras'', and ''Williamsoceras'', have a longitudinal dorsal or ventral process, or both about which the endocones are ...
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