Ellesmeroceratidae
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Ellesmeroceratidae
The Ellesmeroceratidae constitute a family within the cephalopod order Ellesmerocerida. They lived from the Upper Cambrian to the Lower Ordovician. They are characterized by straight and endogastric shells, often laterally compressed, so the dorso-ventral dimension is slightly greater than the lateral, with close spaced sutures having shallow lateral lobes and a generally large tubular ventro-marginal siphuncle with concave segments and irregularly spaced diaphragms. Connecting rings are thick and layered, externally straight but thickening inwardly with the maximum near the middle of the segment so as to leave concave depressions on internal siphuncle molds. Septal necks are typically orthochoanitic but vary in length from almost absent (achoanitic) to reaching halfway to the previous septum (hemichoanitic) and may even slope inwardly (loxochoanitic). Evolution and phylogeny The Ellesmeroceratidae have their derivation in the Plectronoceratidae, order Plectronocerida, in Tr ...
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Ellesmerocerida
The Ellesmerocerida is an order of primitive cephalopods belonging to the subclass Nautiloidea with a widespread distribution that lived during the Late Cambrian and Ordovician. Morphology The Ellesmerocerida are characterized by shells that are typically small, some even tiny, with close-spaced septa and relatively large ventral siphuncles. In some genera (e.g. '' Paleoceras''), the septa are uniformly spaced. Shells of ellesmerocerids are typically smooth and compressed and vary in form. They may be breviconic (short) or longiconic (elongate), straight (orthoconic) or curved (cyrtoconic). Cyrtoconic forms are usually endogastric, with longitudinally convex ventral margins. The apeces of straight forms typically have an endogastric curvature. Some may have grown to as much as 15 cm. Siphuncle segments are tubular or concave. Septal necks are short. Connecting rings which may appear layered are thick and typically wedge shaped with their maximum width at or near where ...
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Ectenolites
''Ectenolites'' is a genus of small, slender, cylindrical Ellesmeroceratids that resemble '' Ellesmeroceras'' but are smaller and proportionally narrower. Septa, as typical for ellesmerocerids, are close spaced with shallow lobes on either flank. The body chamber is proportionally long, the shell itself slightly compressed. The dorsal side at the beginning of the shell, opposite the aperture and body chamber, is strongly convex so to produce a sense of endogastric curvature with the apex and siphuncle The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and ... aligned.Flower R.H.1964, The Nautiloid Order Ellesmeroceratida (Cephalopoda). Memoir 12, New Mexico Bureau of Mines and Mineral Resources, Socorro, NM The siphuncle in ''Ectenolites'' lies along the ventral side; is tubular, composed ...
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Ellesmeroceras Scheii
''Ellesmeroceras'' is the type genus for the Ellesmeroceratidae, a family of primitive nautiloid cephalopods, that is characterized by its small, generally compressed, gradually expanded, orthoconic shell, found in Lower Ordovician marine sediments. The septa are close spaced and the siphuncle is ventral, about 0.2 the diameter of the shell. Septal necks are typically orthochoanitic (short, straight) but may slant inwardly ( loxochoanitic) or reach halfway to the previous septum ( hemichoanitic). Connecting rings are thick. As common for the Ellesmerocerida, ''Ellesmeroceras'' has diaphragms within the siphuncle tube. The type species, ''Ellesmeroceras scheii'', named by Foeste, 1921, was first found on Ellesmere Island in the Canadian arctic, from whence the genus gets its name. ''Ellesmeroceras'' is one of three straight shelled Ellesmeroceratids, the other two being ''Ectenolites ''Ectenolites'' is a genus of small, slender, cylindrical Ellesmeroceratids that resemble '' E ...
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Ellesmeroceras
''Ellesmeroceras'' is the type genus for the Ellesmeroceratidae, a family of primitive nautiloid cephalopods, that is characterized by its small, generally compressed, gradually expanded, orthoconic shell, found in Lower Ordovician marine sediments. The septa are close spaced and the siphuncle is ventral, about 0.2 the diameter of the shell. Septal necks are typically orthochoanitic (short, straight) but may slant inwardly ( loxochoanitic) or reach halfway to the previous septum ( hemichoanitic). Connecting rings are thick. As common for the Ellesmerocerida, ''Ellesmeroceras'' has diaphragms within the siphuncle tube. The type species, ''Ellesmeroceras scheii'', named by Foeste, 1921, was first found on Ellesmere Island in the Canadian arctic, from whence the genus gets its name. ''Ellesmeroceras'' is one of three straight shelled Ellesmeroceratids, the other two being ''Ectenolites ''Ectenolites'' is a genus of small, slender, cylindrical Ellesmeroceratids that resemble '' E ...
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Gasconadian Stage
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Cambrian
The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Ca ...
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Cambrian First Appearances
The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Cambrian biolo ...
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Prehistoric Nautiloid Families
Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of symbols, marks, and images appears very early among humans, but the earliest known writing systems appeared 5000 years ago. It took thousands of years for writing systems to be widely adopted, with writing spreading to almost all cultures by the 19th century. The end of prehistory therefore came at very different times in different places, and the term is less often used in discussing societies where prehistory ended relatively recently. In the early Bronze Age, Sumer in Mesopotamia, the Indus Valley Civilisation, and ancient Egypt were the first civilizations to develop their own scripts and to keep historical records, with their neighbors following. Most other civilizations reached the end of prehistory during the following Iron Age. T ...
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Orthocerida
Orthocerida is an order of extinct Orthoceratoid cephalopods also known as the Michelinocerida that lived from the Early Ordovician () possibly to the Late Triassic (). A fossil found in the Caucasus suggests they may even have survived until the Early Cretaceous (). They were most common however from the Ordovician to the Devonian. Shell form The shell is usually long, and may be straight ("orthoconic") or gently curved. In life, these animals may have been similar to the modern squid, except for the long shell. The internal structure of the shell consists of concavo-convex chambers linked by a centrally-placed tube called a siphuncle. There is a tendency for the chambers to develop cameral deposits, which were used as ballast to balance the long gas-filled shell. Depending on the family, the siphuncle has orthochoanitic (short and straight) or cyrtochoanitic (outwardly curved) septal necks, which protrude from the septa. The shell surface may be (depending on the specie ...
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Tarphycerida
The Tarphycerida were the first of the coiled cephalopods, found in marine sediments from the Lower Ordovician (middle and upper Canad) to the Middle Devonian. Some, such as '' Aphetoceras'' and '' Estonioceras'', are loosely coiled and gyroconic; others, such as '' Campbelloceras'', '' Tarphyceras'', and '' Trocholites'', are tightly coiled, but evolute with all whorls showing. The body chamber of tarphycerids is typically long and tubular,Furnish and Glenister 1964; Nautiloidea - Tarphycerida; In the ''Treatise on Invertebrate Paleontology'' Vol K; Teichert and Moore, (eds) GSA and U of Kansas Press 1964 as much as half the length of the containing whorl in most, greater than in the Silurian Ophidioceratidae. The Tarphycerida evolved from the elongated, compressed, exogastric Bassleroceratidae, probably ''Bassleroceras'', around the end of the Gasconadian through forms like ''Aphetoceras''. Close coiling developed rather quickly, and both gyroconic and evolute forms are fou ...
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Endocerida
Endocerida is an extinct nautiloid order, a group of cephalopods from the Lower Paleozoic with cone-like deposits in their siphuncle. Endocerida was a diverse group of cephalopods that lived from the Early Ordovician possibly to the Late Silurian. Their shells were variable in form. Some were straight (orthoconic), others curved (cyrtoconic); some were long (longiconic), others short (breviconic). Some long-shelled forms like '' Endoceras'' attained shell lengths close to . The related ''Cameroceras'' is anecdotally reported to have reached lengths approaching , but these claims are problematic. The overwhelming majority of endocerids and nautiloids in general are much smaller, usually less than a meter long when fully grown. Morphology Endocerids had a relatively small body chamber as well as a proportionally large siphuncle, which in some genera reached nearly half the shell diameter. This suggests that much of the visceral mass may have been housed within the siphuncle ...
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Clarkoceras
''Clarkoceras'' is a genus of breviconic ellesmerocerid cephalopods, one of only two genera known to have crossed from the Late Cambrian, Trempealeauan, into the Early Ordovician, Gasconadian. (Flower 1964, Teichert 1988); the other being '' Ectenolites''. Description Clarkoceras has a rapidly expanding, laterally compressed, relatively short, endogastrically cyrtoconic shell; The upper or dorsal side is more strongly convex longitudinally than the lower or ventral side is concave. Sutures are essentially straight and close spaced indicating very short camerae (chambers). The siphuncle is relatively large, 0.3 the dorsoventral dimension and is ventral, although not necessarily marginal. (Flower 1964, Furnish and Glenister 1964). Septal necks are straight, reaching about half way back to the previous septum and the connecting rings are thick and layered Taxonomy ''Clarkoceras'' was first thought by Clarke to be a pilocerid, who in 1897 give it the name ''Piloceras newton-winche ...
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