Elanodactylus Prolatus
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Elanodactylus Prolatus
''Elanodactylus'' (meaning "Kite finger") is a genus of ctenochasmatid pterodactyloid pterosaur from the Early Cretaceous period (early Aptian stage) of what is now the Yixian Formation of Liaoning, China. The genus was named in 2008 by Brian Andres and Ji Qiang. The type species is ''Elanodactylus prolatus''. The generic name is derived from the Kite genus ''Elanus'', in reference to the long wings, and Greek ''daktylos'', "finger", referring to the wing finger of pterosaurs. The specific name means "elongated" in Latin, in reference to the elongated middle cervical vertebrae. It is known from a partial postcranial skeleton, holotype NGMC 99-07-1, preserving the wings, sternum, shoulder girdle, ribs, cervical and dorsal vertebrae. It represents an adult individual. In 2010, a second adult was described, specimen LPM-R00078, also a skeleton lacking the skull. The wingspan of the holotype was about , and it is estimated to have weighed . The first phalanx of the wing finger i ...
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Early Cretaceous
The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian-Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining early Aptian Oceanic Anoxic Event 1a (OAE1a) was carbon isotope dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java-Manihiki-Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Ontong Java Plateau today covers an area of 1,860,000 km2. In the Indian Ocean another LIP began to form at c. 120 Ma, the Kerguelen P ...
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Holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering. A holotype is not necessarily "typ ...
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Pterodaustro
''Pterodaustro'' is a genus of ctenochasmatid pterodactyloid pterosaur from South America. Its fossil remains dated back to the Early Cretaceous period, about 105 million years ago. The most distinctive characteristic that separates ''Pterodaustro'' from other ctenochasmatids is its bristle-like teeth, a feature not seen in any other pterosaur. Discovery and naming The first fossils, among them the holotype PVL 2571, a thigh bone, were in the late sixties discovered by Bonaparte in the Lagarcito Formation, situated in the San Luis Province of Argentina, and dating from the Albian. The genus has later also been found in Chile in the Santa Ana Formation. At the Argentine site, the just large "Loma del ''Pterodaustro''", since then, during several expeditions, over 750 ''Pterodaustro'' specimens have been collected, 288 of them having been catalogued until 2008. This makes the species one of the best known pterosaurs, with examples from all growth stages, from egg to adult. Th ...
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Pterodaustrini
Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early Cretaceous periods. The earliest known ctenochasmatid remains date to the Late Jurassic Kimmeridgian age. Previously, a fossil jaw recovered from the Middle Jurassic Stonesfield Slate formation in the United Kingdom, was considered the oldest known. This specimen supposedly represented a member of the family Ctenochasmatidae,Buffetaut, E. and Jeffrey, P. (2012). "A ctenochasmatid pterosaur from the Stonesfield Slate (Bathonian, Middle Jurassic) of Oxfordshire, England." ''Geological Magazine'', (advance online publication) though further examination suggested it actually belonged to a teleosaurid stem-crocodilian instead of a pterosaur. Classification Below is cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "char ...
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Ctenochasma
''Ctenochasma'' (meaning "comb jaw") is a genus of Late Jurassic ctenochasmatid pterosaur belonging to the suborder Pterodactyloidea. Three species are currently recognized: ''C. roemeri'' (named after Friedrich Adolph Roemer), ''C. taqueti'', and ''C. elegans''. Their fossilized remains have been found in the Solnhofen Limestone of Bavaria, Germany, the "Purbeck Group" of northeastern Germany, and the Calcaires tâchetés of eastern France. History The name ''Ctenochasma'' was coined by the German paleontologist Christian Erich Hermann von Meyer in 1852, based on a single lower jaw full of closely packed teeth which he gave the species name ''Ctenochasma roemeri''.von Meyer, C.E.H. (1852). "''Ctenochasma Roemeri''." ''Paläontographica'', 2: 82–84 & pl. 13. A second species, ''C. gracile'', was named by Oppel in 1862 based on a fragmentary skull.Oppel A. (1862). "Über Fährten im lithographischen Schiefer. Paläontologische Mitteilungen aus dem Museum des Koenigl." Bayris ...
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Ctenochasmatinae
Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early Cretaceous periods. The earliest known ctenochasmatid remains date to the Late Jurassic Kimmeridgian age. Previously, a fossil jaw recovered from the Middle Jurassic Stonesfield Slate formation in the United Kingdom, was considered the oldest known. This specimen supposedly represented a member of the family Ctenochasmatidae,Buffetaut, E. and Jeffrey, P. (2012). "A ctenochasmatid pterosaur from the Stonesfield Slate (Bathonian, Middle Jurassic) of Oxfordshire, England." ''Geological Magazine'', (advance online publication) though further examination suggested it actually belonged to a teleosaurid stem-crocodilian instead of a pterosaur. Classification Below is cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "char ...
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Ctenochasmatidae
Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early Cretaceous periods. The earliest known ctenochasmatid remains date to the Late Jurassic Kimmeridgian age. Previously, a fossil jaw recovered from the Middle Jurassic Stonesfield Slate formation in the United Kingdom, was considered the oldest known. This specimen supposedly represented a member of the family Ctenochasmatidae,Buffetaut, E. and Jeffrey, P. (2012). "A ctenochasmatid pterosaur from the Stonesfield Slate (Bathonian, Middle Jurassic) of Oxfordshire, England." ''Geological Magazine'', (advance online publication) though further examination suggested it actually belonged to a teleosaurid stem-crocodilian instead of a pterosaur. Classification Below is cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "cha ...
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Phylogenetic Analysis
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does n ...
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Convergent Evolution
Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last common ancestor of those groups. The cladistic term for the same phenomenon is homoplasy. The recurrent evolution of flight is a classic example, as flying insects, birds, pterosaurs, and bats have independently evolved the useful capacity of flight. Functionally similar features that have arisen through convergent evolution are ''analogous'', whereas '' homologous'' structures or traits have a common origin but can have dissimilar functions. Bird, bat, and pterosaur wings are analogous structures, but their forelimbs are homologous, sharing an ancestral state despite serving different functions. The opposite of convergence is divergent evolution, where related species evolve different traits. Convergent evolution is similar to parallel evo ...
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Cladistics
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived characteristics ( synapomorphies'')'' that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms ''worms'' or ''fishes'' were used within a ''strict'' cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically, outside of cladistics, e.g. as a 'grade', which are fruitless to precisely delineate, especially when including extinct species. R ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and Ant ...
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Azhdarchidae
Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cretaceous as well (late Berriasian age, about 140 million years ago). Azhdarchids included some of the largest known flying animals of all time, but smaller cat-size members have also been found. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the Azhdarchinae to include the pterosaurs ''Azhdarcho'', ''Quetzalcoatlus'', and ''Titanopteryx'' (now known as ''Arambourgiania''). They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. By the time of the end-Cretaceous mass extinction, most pterosaur families except for the Azhdarchidae disappear from the fossil record, but recent studies indicate a wealth of pterosaurian fauna, including pteran ...
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