Ediacaran Orogenies
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Ediacaran Orogenies
The Ediacaran Period ( ) is a geological period that spans 96 million years from the end of the Cryogenian Period 635 million years ago (Mya), to the beginning of the Cambrian Period 538.8 Mya. It marks the end of the Proterozoic Eon, and the beginning of the Phanerozoic Eon. It is named after the Ediacara Hills of South Australia. The Ediacaran Period's status as an official geological period was ratified in 2004 by the International Union of Geological Sciences (IUGS), making it the first new geological period declared in 120 years. Although the period takes its name from the Ediacara Hills where geologist Reg Sprigg first discovered fossils of the eponymous Ediacaran biota in 1946, the type section is located in the bed of the Enorama Creek within Brachina Gorge in the Flinders Ranges of South Australia, at . The Ediacaran marks the first appearance of widespread multicellular fauna following the end of Snowball Earth glaciation events, the so-called Ediacaran biota, wh ...
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International Commission On Stratigraphy
The International Commission on Stratigraphy (ICS), sometimes referred to unofficially as the "International Stratigraphic Commission", is a daughter or major subcommittee grade scientific daughter organization that concerns itself with stratigraphy, stratigraphical, geology, geological, and chronology, geochronological matters on a global scale. It is the largest subordinate body of the International Union of Geological Sciences (IUGS). The ICS is essentially a permanent working committee, working subcommittee, which meets far more regularly than the quadrennial meetings scheduled by the IUGS, when it meets as a congress or committee, membership of the whole. Aims One of its main aims, a project begun in 1974, is to establish a multidisciplinary standard and global geologic time scale that will ease paleontology, paleontological and geobiology, geobiological comparisons region to region by benchmarks with stringent and rigorous strata criteria called Global Boundary Stratotype ...
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Ediacaran Biota
The Ediacaran (; formerly Vendian) biota is a Taxonomy (biology), taxonomic period classification that consists of all life forms that were present on Earth during the Ediacaran Period (). These were composed of enigmatic tubular and frond-shaped, mostly sessility (zoology), sessile, organisms. Trace fossils of these organisms have been found worldwide, and represent the earliest known complex multicellular organisms. The Ediacaran biota may have undergone evolutionary radiation in a proposed event called the Avalon explosion, . This was after the Earth had thawed from the Cryogenian period's Snowball Earth, extensive glaciation. This biota largely disappeared with the rapid increase in biodiversity known as the Cambrian explosion. Most of the currently existing body plans of animals first appeared in the fossil, fossil record of the Cambrian rather than the Ediacaran. For macroorganisms, the Cambrian biota appears to have almost completely replaced the organisms that dominate ...
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Tribrachidium
''Tribrachidium heraldicum'' is a tri-radially symmetric fossil animal that lived in the late Ediacaran (Vendian) seas. In life, it was hemispherical in form. ''T. heraldicum'' is the best known member of the extinct group Trilobozoa. Etymology The generic name ''Tribrachidium'' is derived from combination of the grc, τρία (, "three") + la, brachium ("arm") + diminutive suffix . The specific name ''T. heraldicum'' references the similarity of the pattern of this fossil with the well-known heraldic triskelion design, such as the coat of arms of the Isle of Man. Occurrence ''Tribrachidium'' fossils were first discovered in the Ediacara Member of the Rawnslay Quartzite, Flinders Ranges in South Australia. This fossil is also known from the Mogilev Formation in the Dniester River Basin, Podolia, Ukraine and from the Verkhovka, Zimnegory and Yorga Formations in the White Sea area of the Arkhangelsk Region, Russia. Description ''T. heraldicum'' is preserved as negative impressio ...
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Trilobozoa
''Trilobozoa'' (meaning "three-lobed animals") is a phylum of extinct mobile animals that were originally classified into the Cnidaria. The basic body plan of ''Trilobozoa'' is often a tri-radial or radial sphere-shaped form with lobes radiating from its centre. Fossils of trilobozoans are restricted to marine strata of the Late Ediacaran period. History and interpretations Originally, both Fedonkin and Runnegar presumed that there were 2-3 families within the Trilobozoa, those families being Albumaresidae Fedonkin, 1985 and Tribrachididae Runnegar, 1992. Although, affinities with the Conulariida were made because of the conulariids possessing similar three-fold symmetry. Mikhail Fedonkin later classified the Trilobozoa as a class of the Phylum Coelenterata. Most of the members of what is now the modern day classification for Trilobozoa were thought to have originally been free swimming Jellyfish. Tribrachidium was once interpreted as a Edrioasteroid Echinoderm, although wi ...
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Cyclomedusa
''Cyclomedusa'' is a circular fossil of the Ediacaran biota; it has a circular bump in the middle and as many as five circular growth ridges around it. Many specimens are small, but specimens in excess of 20 cm are known. The concentric disks are not necessarily circular, especially when adjacent individuals interfere with each other's growth. Many radial segment lines — somewhat pineapple-like — extend across the outer disks. A few specimens show what might be a stem extending from the center in some direction or other. ''Cyclomedusa'' is widely distributed in Ediacaran strata, with a number of species described. It has also been found in sediments dating to the Mayanian (~). ''Cyclomedusa'' was originally thought to be a jellyfish but some specimens seem to be distorted to accommodate adjacent specimens on the substrate, apparently indicating a benthic (bottom-dwelling) creature. The markings do not match the musculature pattern of modern jellyfish. The fossils hav ...
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Aspidella
:Aspidella'' is also a homonym for the mushroom genus ''Saproamanita. ''Aspidella'' is an Ediacaran disk-shaped fossil of uncertain affinity. It is known from the single species ''A. terranovica''. Morphology ''Aspidella'' consists of disk-shaped fossils, with concentric rings and/or centripetal rays. The diameter of circular ''Aspidella'' varies from 1 to 180 mm.Peterson. P. 131 Most individuals are between 4 and 10 mm, but smaller individuals would presumably have decayed before they could fossilize. Other ''Aspidella'' take the form of ellipses, 3–8 cm long and 1–4 cm wide. Most have a central pimple. The rim of all specimens is made up by ridge-edged rays and/or concentric rings. Ecology The rarity of large individuals probably indicates that ''Aspidella'' were r-strategists, producing numerous offspring of which most died young. It is most common in deep-water sediments, but is a constituent of most Ediacaran fossil assemblages, including tho ...
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Charnia
''Charnia'' is a genus of frond-like lifeforms belonging to the Ediacaran biota with segmented, leaf-like ridges branching alternately to the right and left from a zig-zag medial suture (thus exhibiting glide reflection, or opposite isometry). The genus ''Charnia'' was named after Charnwood Forest in Leicestershire, England, where the first fossilised specimen was found. ''Charnia'' is significant because it was the first Precambrian fossil to be recognized as such. The living organism grew on the sea floor and is believed to have fed on nutrients in the water. Despite ''Charnia'' fern-like appearance, it is not a photosynthetic plant or alga because the nature of the fossilbeds where specimens have been found implies that it originally lived in deep water, well below the photic zone where photosynthesis can occur. Diversity Several ''Charnia'' species were described but only the type species ''C. masoni'' is considered valid. Some specimens of ''C. masoni'' were described as ...
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Sea Pen
Sea pens are colonial marine cnidarians belonging to the order Pennatulacea. There are 14 families within the order; 35 extant genera, and it is estimated that of 450 described species, around 200 are valid. Sea pens have a cosmopolitan distribution, being found in tropical and temperate waters worldwide, as well as from the intertidal to depths of more than 6100 m. Sea pens are grouped with the octocorals, together with sea whips ('' gorgonians''). Although the group is named for its supposed resemblance to antique quill pens, only sea pen species belonging to the suborder Subselliflorae live up to the comparison. Those belonging to the much larger suborder Sessiliflorae lack feathery structures and grow in club-like or radiating forms. The latter suborder includes what are commonly known as sea pansies. The earliest accepted fossils are known from the Cambrian-aged Burgess Shale (''Thaumaptilon''). Similar fossils from the Ediacaran (ala ''Charnia'') may ...
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Petalonamae
The petalonamids (Petalonamae) are an extinct group of archaic animals typical of the Ediacaran biota, also called frondomorphs, dating from approximately 635 million years ago to 516 million years ago. They are benthic and motionless animals, that have the shape of leaves, fronds (frondomorphic), feathers or spindles and were initially considered algae, octocorals or sea pens. It is now believed that there are no living descendants of the group, which shares a probable relation to the Ediacaran animals known as Vendozoans. It is considered that the organisms were fluffy at least in appearance, as if "inflatable." They are particularly difficult to classify phylogenetically. Lacking mouths, intestines, reproductive organs, and with no preserved evidence of internal structures, these organisms' existence is very strange by current standards. The most widely accepted hypothesis is that they could suck nutrients from the water around them by osmosis. The fronds were composed of bra ...
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Spriggina
''Spriggina'' is a genus of early bilaterian animals whose relationship to living animals is unclear. Fossils of ''Spriggina'' are known from the late Ediacaran period in what is now South Australia. ''Spriggina floundersi'' is the official fossil emblem of South Australia. It has been found nowhere else. The organism reached about in length and may have been predatory. Its bottom was covered with two rows of tough interlocking plates, while one row covered its top; its front few segments fused to form a "head." ''Spriggina'' affinity is currently unknown; it has been variously classified as an annelid worm, a rangeomorph-like frond, a variant of ''Charniodiscus'', a proarticulatan, or an arthropod perhaps related to the trilobites, or even an extinct phylum. Lack of known segmented legs or limbs, and glide reflection instead of symmetric segments, suggest an arthropod classification is unlikely despite some superficial resemblance. The genus ''Spriggina'' may have originally co ...
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Dickinsonia
''Dickinsonia'' is an extinct genus of basal animal that lived during the late Ediacaran period in what is now Australia, China, Russia and Ukraine. The individual ''Dickinsonia'' typically resembles a bilaterally symmetrical ribbed oval. Its affinities are presently unknown; its mode of growth is consistent with a stem-group bilaterian affinity, though some have suggested that it belongs to the fungi, or even an "extinct kingdom". It lived during the late Ediacaran (part of Precambrian). The discovery of cholesterol molecules in fossils of ''Dickinsonia'' lends support to the idea that ''Dickinsonia'' was an animal. Description ''Dickinsonia'' fossils are known only in the form of imprints and casts in sandstone beds. The specimens found range from a few millimetres to about in length, and from a fraction of a millimetre to a few millimetres thick. They are nearly bilaterally symmetric, segmented, round or oval in outline, slightly expanded to one end (i.e. egg-shaped out ...
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Proarticulata
Proarticulata is a proposed phylum of extinct, bilaterally symmetrical animals known from fossils found in the Ediacaran (Vendian) marine deposits, and dates to approximately . The name comes from the Greek () = "before" and Articulata, i.e. prior to animals with true segmentation such as annelids and arthropods. This phylum was established by Mikhail A. Fedonkin in 1985 for such animals as '' Dickinsonia'', ''Vendia'', ''Cephalonega'', ''Praecambridium'' and currently many other Proarticulata are described (see list). Due to their simplistic morphology, their affinities and mode of life are subject to debate. They are almost universally considered to be metazoans, and due to possessing a clear central axis have been suggested to be stem-bilaterians. In the traditional interpretation, the Proarticulatan body is divided into transverse articulation (division) into isomers as distinct from the transverse articulation segments in annelids and arthropods, as their individual isome ...
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