Dsungaripteromorpha
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Dsungaripteromorpha
Azhdarchoidea (or azhdarchoids) is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea. Pterosaurs belonging to this group lived throughout the Early and Late Cretaceous periods, with one tentative member, ''Tendaguripterus'', that lived in the Late Jurassic period. The largest azhdarchoids include members of the family Azhdarchidae, examples of these are ''Quetzalcoatlus'', '' Hatzegopteryx'', and ''Arambourgiania''. The Azhdarchoidea has been recovered as either closely related to the Ctenochasmatoidea, as the sister taxon of the Pteranodontoidea within the Ornithocheiroidea, or within the Tapejaroidea, which in turn was also within the Ornithocheiroidea. Classification Azhdarchoidea was given a phylogenetic definition by David Unwin in 2003. Unwin defined the group as the most recent common ancestor of ''Quetzalcoatlus'' and '' Tapejara'', and all its descendants.Unwin, D. M., (2003). "On the phylogeny and evolutio ...
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Eoazhdarcho
''Eoazhdarcho'' is a genus of azhdarchoid pterodactyloid pterosaur named in 2005 by Chinese paleontologists Lü Junchang and Ji Qiang. The type and only known species is ''Eoazhdarcho liaoxiensis''. The fossil was found in the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. Etymology The genus name, ''Eoazhdarcho'', combines a Greek ''eos'', "dawn" with the name of the genus ''Azhdarcho'', with the implication it was an early related form of the latter. The specific name, ''liaoxiensis'', refers to the ancient region Liaoxi. Description ''Eoazhdarcho'' is based on holotype GMN-03-11-02, a partial skeleton and lower jaw, and is distinguished from other pterosaurs by the proportions of its bones. The metacarpals are very elongated but the cervical vertebrae and hind limbs are not. It was relatively small by azhdarchoid standards, with a wingspan of about 1.6 meters (5.2 feet). Classification The describers first assigned ''Eoazhdarcho'' t ...
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Pterodactyloidea
Pterodactyloidea (derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional suborders of pterosaurs ("wing lizards"), and contains the most derived members of this group of flying reptiles. They appeared during the middle Jurassic Period, and differ from the basal (though paraphyletic) rhamphorhynchoids by their short tails and long wing metacarpals (hand bones). The most advanced forms also lack teeth, and by the late Cretaceous, all known pterodactyloids were toothless. Many species had well-developed crests on the skull, a form of display taken to extremes in giant-crested forms like ''Nyctosaurus'' and ''Tupandactylus''. Pterodactyloids were the last surviving pterosaurs when the order became extinct at the end of the Cretaceous Period, together with the non-avian dinosaurs and most marine reptiles. "Pteroda ...
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Dsungaripteridae
Dsungaripteridae is a group of pterosaurs within the suborder Pterodactyloidea. They were robust pterosaurs with good terrestrial abilities and flight honed for inland settings. Classification In 1964 Young created a family to place the recently found Chinese genus ''Dsungaripterus''. Later on, also ''Noripterus'' (then now with the name "Phobetor" which was already occupied, therefore the quotation marks) were assigned to the family. In 2003, Alexander Kellner gave the exact definition as a clade:Kellner, A.W.A., 2003. Pterosaur phylogeny and comments on the evolutionary history of the AN group. In: Buffetaut, E., Mazin, J.M. (Eds.), ''Evolution and Palaeobiology of Pterosaurs. Geological Society, London, Special Publication'' 217, 105–137. the group was composed out of the latest common ancestor of ''Dsungaripterus'', ''Noripterus'' and “Phobetor”, and all its descendants. As synapomorphies he gave the next six characteristics: a relatively small eye-socket, which is pla ...
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Radiodactylus
''Radiodactylus'' (meaning "radio finger") is an extinct genus of non-azhdarchid azhdarchoid pterosaur known from the Early Cretaceous period of what is now Texas, southern United States. It contains a single species, ''Radiodactylus langstoni''. Discovery ''Radiodactylus'' is known solely from its holotype, SMU 72547, a nearly complete left humerus first described by Phillip Murry ''et al.'' (1991).Murry, P.A., Winkler, D.A. & Jacobs, L.L., 1991, "An azhdarchid pterosaur humerus from the Lower Cretaceous Glen Rose Formation of Texas", ''Journal of Paleontology'' 65(1): 167–170 The specimen is well preserved in three dimensions with no apparent crushing. The humerus lacks only portions of the proximal end and anterior end of the deltopectoral crest, and has a fracture in the mid-shaft area where it is very slightly rotationally distorted. ''Radiodactylus'' was first named by Brian Andres and Timothy S. Myers in 2013 and the type species is ''Radiodactylus langstoni''. The ...
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Alanqa
''Alanqa'' is a genus of pterodactyloid pterosaur from the Late Cretaceous period ( Cenomanian stage) of what is now the Kem Kem Beds of southeastern Morocco. The name ''Alanqa'' comes from the Arabic word العنقاء ''al- ‘Anqā’'', for a mythical bird of Arabian culture. Discovery Aided by local villagers, a team of paleontologists had been excavating at several locations in the Kem Kem Beds during April, and November to December 2008, uncovering remains of several different pterosaurs. The material was fragmentary, and the type locality for ''Alanqa'' is Aferdou N'Chaft, near the village of Begaa and to the north-east of Taouz. Description ''Alanqa'' is known only from five fragments of the front upper and lower jaws, and possibly a neck vertebra, representing the single type species ''Alanqa saharica''. Two of these fragments were first described, but not named, by Wellnhofer and Buffetaut in 1999, and referred to a pteranodontid. Three additional jaw specimens, ...
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Ornithocheiroidea
Ornithocheiroidea (or ornithocheiroids) is a group of pterosaurs within the extinct suborder Pterodactyloidea. They were typically large pterosaurs that lived from the Early to Late Cretaceous periods (Valanginian to Maastrichtian stages), with fossil remains found all over the world except Antarctica. Ornithocheiroids were the most advanced group of pterosaurs, as the group includes the clade Azhdarchoidea, of which its members lived until the Maastrichtian stage of the Late Cretaceous, around 66 million years ago. Notable pterosaurs from this group include the pteranodontians ''Pteranodon'' and ''Nyctosaurus'', the ornithocheirid ''Ornithocheirus'', the anhanguerid ''Tropeognathus'', as well as the azhdarchids ''Hatzegopteryx'' and ''Quetzalcoatlus''. Classification The name Ornithocheiroidea was originally defined as an apomorphy-based taxon by Christopher Bennett in 1994. It was given a relationship-based definition in 2003 by Alexander Kellner, who defined it as the least in ...
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Microtuban
''Microtuban'' is an extinct genus of azhdarchoid pterosaur known from the Late Cretaceous of northern Lebanon. Discovery ''Microtuban'' is known only from a partially preserved skeleton lacking the skull, holotype SMNK PAL 6595. It was acquired from a local fossil dealer by the ''Staatliches Museum für Naturkunde Karlsruhe'' and was, as can be deduced from the qualities of the stone matrix, collected in the locality of Hjoûla in a marine layer of the Sannine Formation, dating to the early Cenomanian stage of the Late Cretaceous, about 99.6–96 million years ago. The fossil dealer however, indicated it came from a quarry at nearby Hâqel. Regardless, it is only the second pterosaur fossil found in Lebanon and a rare example of a pterosaur from the African continental plate, to which the area in Cretaceous times belonged. In this period the site was located hundreds of miles from the mainland coastline. The specimen consists of the last cervical and first dorsal vertebrae, a ...
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Pterosaurs
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insectiv ...
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Quetzalcoatlus
''Quetzalcoatlus'' is a genus of pterosaur known from the Late Cretaceous period of North America (Maastrichtian stage); its members were among the largest known flying animals of all time. ''Quetzalcoatlus'' is a member of the Azhdarchidae, a family of advanced toothless pterosaurs with unusually long, stiffened necks. Its name comes from the Aztec feathered serpent god Quetzalcoatl. The type species is ''Q. northropi'', named by Douglas Lawson in 1975; the genus also includes the smaller species ''Q. lawsoni'', which was known for many years as an unnamed species before being named by Brian Andres and Wann Langston Jr. (posthumously) in 2021. Discovery and species The first ''Quetzalcoatlus'' fossils were discovered in Texas, United States, from the Maastrichtian Javelina Formation at Big Bend National Park (dated to around 68 million years ago) in 1971 by Douglas A. Lawson, then a geology graduate student from the Jackson School of Geosciences at the University of Texas ...
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Tendaguripterus
''Tendaguripterus'' was a genus of pterodactyloid pterosaur from the Kimmeridgian to Tithonian-age Upper Jurassic Middle Saurian Beds (Tendaguru Formation) of Tendaguru, Lindi Region, Tanzania. Discovery and naming During the German paleontological expeditions to German East Africa between 1909 and 1913, some pterosaur fossil material was collected that was recognized as such by Hans Reck in 1931. In 1999 David Unwin and Wolf-Dieter Heinrich named a new genus for it. The type species is ''Tendaguripterus recki''. The genus name is derived from Tendaguru and a Latinized Greek ''pteron'', "wing". The specific name honors Reck. Description The genus is based on holotype MB.R.1290, a partial mandible with teeth (the symphyseal region, where the two lower jaws meet and fuse into one element). The top of the back of the symphysis is very concave. The teeth in the posterior section of the jaw fragment point strongly backwards. They are also the longest. The teeth are set relativel ...
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Hatzegopteryx
''Hatzegopteryx'' ("Hațeg basin wing") is a genus of azhdarchid pterosaur found in the late Maastrichtian deposits of the Densuş Ciula Formation, an outcropping in Transylvania, Romania. It is known only from the type species, ''Hatzegopteryx thambema'', named by Buffetaut ''et al.'' in 2002 based on parts of the skull and humerus. Additional specimens, including a neck vertebra, were later placed in the genus, representing a range of sizes. The largest of these remains indicate it was among the biggest pterosaurs, with an estimated wingspan of . Unusually among giant azhdarchids, ''Hatzegopteryx'' had a very wide skull bearing large muscular attachments; bones with a spongy internal texture instead of hollow; and a short, robust, and heavily muscled neck measuring long, which was about half the length of other azhdarchids with comparable wingspans, and was capable of withstanding strong bending forces. ''Hatzegopteryx'' inhabited Hațeg Island, an island situated in the Cret ...
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Arambourgiania
''Arambourgiania'' is an extinct genus of azhdarchid pterosaur from the Late Cretaceous period (Maastrichtian stage) of Jordan, and possibly the United States.Harrell, T. Lynn Jr.; Gibson, Michael A.; Langston, Wann Jr. (2016). "A cervical vertebra of ''Arambourgiania philadelphiae'' (Pterosauria, Azhdarchidae) from the Late Campanian micaceous facies of the Coon Creek Formation in McNairy County, Tennessee, USA" ''Bull. Alabama Mus. Nat. Hist.'' 33:94–103 ''Arambourgiania'' was among the largest members of its family, the Azhdarchidae, and it is also one of the largest flying animals ever known. The incomplete left ulna of the "Sidi Chennane azhdarchid" from Morocco may have also belonged to ''Arambourgiania''. History of discovery In the early 1940s, a railway worker during repairs on the Amman-Damascus railroad near Russeifa found a two foot long fossil bone. In 1943 this was acquired by the director of a nearby phosphate mine, Amin Kawar, who brought it to the attention ...
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