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Discredited Hypotheses For The Cambrian Explosion
As understanding of the events of the Cambrian becomes clearer, data have accumulated to make some postulated causes for the Cambrian explosion look improbable. Some examples are the evolution of herbivory, vast changes in plate tectonic rates or orbital motion, or different evolutionary mechanisms in force. Developmental explanations Regulatory genes Hox genes regulate the operation of other genes by switching them on or off in various parts of the body, for example “make an eye here” or “make a leg there”. Very similar Hox genes are found in all animals from Cnidaria (e.g. jellyfish) to humans, although mammals have four sets of Hox genes while Cnidaria have only one. Hox genes in different animal groups are so similar that, for example, one can transplant a human “make an eye” Pax6 gene into a fruitfly embryo and it still causes an eye to form – but it is a fruitfly eye, because the genes that the transplanted Hox gene activates are fruitfly genes. The fact tha ...
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Cambrian Explosion
The Cambrian explosion, Cambrian radiation, Cambrian diversification, or the Biological Big Bang refers to an interval of time approximately in the Cambrian Period when practically all major animal phyla started appearing in the fossil record. It lasted for about 13 – 25 million years and resulted in the divergence of most modern metazoan phyla. The event was accompanied by major diversification in other groups of organisms as well. Before early Cambrian diversification, most organisms were relatively simple, composed of individual cells, or small multicellular organisms, occasionally organized into colonies. As the rate of diversification subsequently accelerated, the variety of life became much more complex, and began to resemble that of today. Almost all present-day animal phyla appeared during this period, including the earliest chordates. A 2019 paper suggests that the timing should be expanded back to include the late Ediacaran, rather than just the n ...
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Plate Tectonic
Plate tectonics (from the la, label=Late Latin, tectonicus, from the grc, τεκτονικός, lit=pertaining to building) is the generally accepted scientific theory that considers the Earth's lithosphere to comprise a number of large tectonic plates which have been slowly moving since about 3.4 billion years ago. The model builds on the concept of ''continental drift'', an idea developed during the first decades of the 20th century. Plate tectonics came to be generally accepted by geoscientists after seafloor spreading was validated in the mid to late 1960s. Earth's lithosphere, which is the rigid outermost shell of the planet (the crust and upper mantle), is broken into seven or eight major plates (depending on how they are defined) and many minor plates or "platelets". Where the plates meet, their relative motion determines the type of plate boundary: '' convergent'', '' divergent'', or ''transform''. Earthquakes, volcanic activity, mountain-building, and oceanic tren ...
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Hox Gene
Hox genes, a subset of homeobox genes, are a group of related genes that specify regions of the body plan of an embryo along the head-tail axis of animals. Hox proteins encode and specify the characteristics of 'position', ensuring that the correct structures form in the correct places of the body. For example, Hox genes in insects specify which appendages form on a segment (for example, legs, antennae, and wings in fruit flies), and Hox genes in vertebrates specify the types and shape of vertebrae that will form. In segmented animals, Hox proteins thus confer segmental or positional identity, but do not form the actual segments themselves. Studies on Hox genes in ciliated larvae have shown they are only expressed in future adult tissues. In larvae with gradual metamorphosis the Hox genes are activated in tissues of the larval body, generally in the trunk region, that will be maintained through metamorphosis. In larvae with complete metamorphosis the Hox genes are mainly express ...
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Cnidaria
Cnidaria () is a phylum under kingdom Animalia containing over 11,000 species of aquatic animals found both in freshwater and marine environments, predominantly the latter. Their distinguishing feature is cnidocytes, specialized cells that they use mainly for capturing prey. Their bodies consist of mesoglea, a non-living jelly-like substance, sandwiched between two layers of epithelium that are mostly one cell thick. Cnidarians mostly have two basic body forms: swimming medusae and sessile polyps, both of which are radially symmetrical with mouths surrounded by tentacles that bear cnidocytes. Both forms have a single orifice and body cavity that are used for digestion and respiration. Many cnidarian species produce colonies that are single organisms composed of medusa-like or polyp-like zooids, or both (hence they are trimorphic). Cnidarians' activities are coordinated by a decentralized nerve net and simple receptors. Several free-swimming species of Cubozoa and Scyphozo ...
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Pax6
Paired box protein Pax-6, also known as aniridia type II protein (AN2) or oculorhombin, is a protein that in humans is encoded by the ''PAX6'' gene. Function PAX6 is a member of the Pax gene family which is responsible for carrying the genetic information that will encode the Pax-6 protein. It acts as a "master control" gene for the development of eyes and other sensory organs, certain neural and epidermal tissues as well as other homologous structures, usually derived from ectodermal tissues. However, it has been recognized that a suite of genes is necessary for eye development, and therefore the term of "master control" gene may be inaccurate. Pax-6 is expressed as a transcription factor when neural ectoderm receives a combination of weak Sonic hedgehog (SHH) and strong TGF-Beta signaling gradients. Expression is first seen in the forebrain, hindbrain, head ectoderm and spinal cord followed by later expression in midbrain. This transcription factor is most noted for its u ...
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Drosophila
''Drosophila'' () is a genus of flies, belonging to the family Drosophilidae, whose members are often called "small fruit flies" or (less frequently) pomace flies, vinegar flies, or wine flies, a reference to the characteristic of many species to linger around overripe or rotting fruit. They should not be confused with the Tephritidae, a related family, which are also called fruit flies (sometimes referred to as "true fruit flies"); tephritids feed primarily on unripe or ripe fruit, with many species being regarded as destructive agricultural pests, especially the Mediterranean fruit fly. One species of ''Drosophila'' in particular, ''D. melanogaster'', has been heavily used in research in genetics and is a common model organism in developmental biology. The terms "fruit fly" and "''Drosophila''" are often used synonymously with ''D. melanogaster'' in modern biological literature. The entire genus, however, contains more than 1,500 species and is very diverse in appearance, be ...
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Embryo
An embryo is an initial stage of development of a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male sperm cell. The resulting fusion of these two cells produces a single-celled zygote that undergoes many cell divisions that produce cells known as blastomeres. The blastomeres are arranged as a solid ball that when reaching a certain size, called a morula, takes in fluid to create a cavity called a blastocoel. The structure is then termed a blastula, or a blastocyst in mammals. The mammalian blastocyst hatches before implantating into the endometrial lining of the womb. Once implanted the embryo will continue its development through the next stages of gastrulation, neurulation, and organogenesis. Gastrulation is the formation of the three germ layers that will form all of the different parts of the body. Neurulation forms the nervous ...
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Compound Eye
A compound eye is a visual organ found in arthropods such as insects and crustaceans. It may consist of thousands of ommatidia, which are tiny independent photoreception units that consist of a cornea, lens, and photoreceptor cells which distinguish brightness and color. The image perceived by this arthropod eye is a combination of inputs from the numerous ommatidia, which are oriented to point in slightly different directions. Compared with single-aperture eyes, compound eyes have poor image resolution; however, they possess a very large view angle and the ability to detect fast movement and, in some cases, the polarization of light. Because a compound eye is made up of a collection of ommatidia, each with its own lens, light will enter each ommatidium instead of using a single entrance point. The individual light receptors behind each lens are then turned on and off due to a series of changes in the light intensity during movement or when an object in moving, creating a flic ...
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Quantum Evolution
Quantum evolution is a component of George Gaylord Simpson's multi-tempoed theory of evolution proposed to explain the rapid emergence of higher taxonomic groups In biology, taxonomy () is the scientific study of naming, defining ( circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon) and these groups are given ... in the fossil record. According to Simpson, Rate of evolution, evolutionary rates differ from group to group and even among closely related lineages. These different rates of evolutionary change were designated by Simpson as ''bradytelic'' (slow tempo), ''horotelic'' (medium tempo), and ''tachytelic'' (rapid tempo). Quantum evolution differed from these styles of change in that it involved a drastic shift in the Evolutionary landscape, adaptive zones of certain classes of animals. The word "Atomic electron transition, quantum" therefore refers to an "all-or-none reaction", ...
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Stanley Awramik
Stanley Awramik (born 1946) is an American biogeologist and paleontologist. He is best known for his work related to the Precambrian. In 2013, he was inducted as a fellow of the Geological Society of America. Career Born in New England, he first studied stromatolites as a graduate student under famed paleobotanist Elso Sterrenberg Barghoorn at Harvard University. In 1979, he became a faculty member at the University of California, Santa Barbara, where he was a contemporary of Preston Cloud. Stromatolites Awramik is known for his long standing expertise on stromatolites. His 1971 discovery of a link between a loss in stromatolitic diversity and the appearance of the first metazoans was published in ''Science''. In 1974, he and Lynn Margulis gave an often-used definition of "stromatolite". Origin of life In 1983, he discovered what was then considered to be the oldest evidence of life on earth, located in Western Australia. His and others' work provided credence to the notion tha ...
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Protists
A protist () is any eukaryotic organism (that is, an organism whose cells contain a cell nucleus) that is not an animal, plant, or fungus. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor), the exclusion of other eukaryotes means that protists do not form a natural group, or clade. Therefore, some protists may be more closely related to animals, plants, or fungi than they are to other protists. However, like the groups ''algae'', ''invertebrates'', and ''protozoans'', the biological category ''protist'' is used for convenience. Others classify any unicellular eukaryotic microorganism as a protist. The study of protists is termed protistology. History The classification of a third kingdom separate from animals and plants was first proposed by John Hogg in 1860 as the kingdom Protoctista; in 1866 Ernst Haeckel also proposed a third kingdom Protista as "the kingdom of primitive forms". Originally these also included prokaryotes, but ...
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Eukaryotes
Eukaryotes () are organisms whose cells have a nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the three domains of life. Bacteria and Archaea (both prokaryotes) make up the other two domains. The eukaryotes are usually now regarded as having emerged in the Archaea or as a sister of the Asgard archaea. This implies that there are only two domains of life, Bacteria and Archaea, with eukaryotes incorporated among archaea. Eukaryotes represent a small minority of the number of organisms, but, due to their generally much larger size, their collective global biomass is estimated to be about equal to that of prokaryotes. Eukaryotes emerged approximately 2.3–1.8 billion years ago, during the Proterozoic eon, likely as flagellated phagotrophs. Their name comes from the Greek εὖ (''eu'', "well" or "good") and κάρυον (''karyon'', "nut" or "kernel"). Eu ...
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