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Dirina Arabica
''Dirina arabica'' is a species of saxicolous (rock-dwelling), crustose lichen in the family Roccellaceae. Found in Socotra, it was formally described as a new species in 2013 by lichenologists Anders Tehler and Damien Ernst. The type specimen was collected by the first author near the village in Homill, at an altitude of . The species epithet refers to Arabia, the geographical location encompassing the type locality. The lichen is endemic to Socotra, where it grows on Eocene limestone rocks. It has a creamy-white, slightly thallus that is 0.1–0.5 mm thick and a chalk-like medulla. There are no soralia on the thallus. The ascomata have a circular outline and measure up to 2.0 mm in diameter, and have a pruinose, white-grey with a . ''Dirina arabica'' is a sister species to '' Dirina immersa'', a sympatric In biology, two related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter one ano ...
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Saxicolous Lichen
A saxicolous lichen is a lichen that grows on rock. The prefix "sax" from the Latin means "rock" or "stone". Characteristics Saxicolous lichens exhibit very slow growth rates. They may develop on rock substrates for long periods of time, given the absence of external disturbances. The importance of the mineral composition of the rock substrate, as well as the elemental geochemistry Geochemistry is the science that uses the tools and principles of chemistry to explain the mechanisms behind major geological systems such as the Earth's crust and its oceans. The realm of geochemistry extends beyond the Earth, encompassing the e ... is also important to the distribution of saxicolous lichens, but the relationship between the substrate influence on lichens, either chemical or textural, is still obscure. Communities of saxicolous lichens are often species-rich in terms of number. References Lichenology {{lichen-stub ...
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Thallus
Thallus (plural: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. Many of these organisms were previously known as the thallophytes, a polyphyletic group of distantly related organisms. An organism or structure resembling a thallus is called thalloid, thallodal, thalliform, thalline, or thallose. A thallus usually names the entire body of a multicellular non-moving organism in which there is no organization of the tissues into organs. Even though thalli do not have organized and distinct parts (leaves, roots, and stems) as do the vascular plants, they may have analogous structures that resemble their vascular "equivalents". The analogous structures have similar function or macroscopic structure, but different microscopic structure; for example, no thallus has vascular tissue. In exceptional cases such as the Lemnoideae, where ...
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Lichen Species
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

Dirina
''Dirina'' is a genus of lichen-forming fungi in the family Roccellaceae. All ''Dirina'' species are crustose lichens with a whitish to greyish brown thallus, and live either on rock or on bark–some species can live on both. The partner is a member of the green algal genus '' Trentepohlia''. Most species occur in the Northern Hemisphere, and are generally restricted to coastal habitats, where they may be locally quite common. Erythrin and lecanoric acid are lichen products that usually occur in ''Dirina'' species, along with several other unidentified substances. The genus was circumscribed in 1825 by Elias Magnus Fries. Swedish lichenologist Anders Tehler published a monograph about the genus in 1983. Thirty years later, he and his colleagues revisited ''Dirina'', combining evidence from molecular phylogenetic analysis with morphological and chemical analysis. They accepted 24 species in ''Dirina'', 9 of which were described as new to science. Species * '' Dirina ango ...
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Sympatry
In biology, two related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter one another. An initially interbreeding population that splits into two or more distinct species sharing a common range exemplifies sympatric speciation. Such speciation may be a product of reproductive isolation – which prevents hybrid offspring from being viable or able to reproduce, thereby reducing gene flow – that results in genetic divergence. Sympatric speciation may, but need not, arise through secondary contact, which refers to speciation or divergence in allopatry followed by range expansions leading to an area of sympatry. Sympatric species or taxa in secondary contact may or may not interbreed. Types of populations Four main types of population pairs exist in nature. Sympatric populations (or species) contrast with parapatric populations, which contact one another in adjacent but not shared ranges and do not ...
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Dirina Immersa
''Dirina'' is a genus of lichen-forming fungi in the family Roccellaceae. All ''Dirina'' species are crustose lichens with a whitish to greyish brown thallus, and live either on rock or on bark–some species can live on both. The partner is a member of the green algal genus '' Trentepohlia''. Most species occur in the Northern Hemisphere, and are generally restricted to coastal habitats, where they may be locally quite common. Erythrin and lecanoric acid are lichen products that usually occur in ''Dirina'' species, along with several other unidentified substances. The genus was circumscribed in 1825 by Elias Magnus Fries. Swedish lichenologist Anders Tehler published a monograph about the genus in 1983. Thirty years later, he and his colleagues revisited ''Dirina'', combining evidence from molecular phylogenetic analysis with morphological and chemical analysis. They accepted 24 species in ''Dirina'', 9 of which were described as new to science. Species * ''Dirina ango ...
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Ascomata
An ascocarp, or ascoma (), is the fruiting body ( sporocarp) of an ascomycete phylum fungus. It consists of very tightly interwoven hyphae and millions of embedded asci, each of which typically contains four to eight ascospores. Ascocarps are most commonly bowl-shaped (apothecia) but may take on a spherical or flask-like form that has a pore opening to release spores (perithecia) or no opening (cleistothecia). Classification The ascocarp is classified according to its placement (in ways not fundamental to the basic taxonomy). It is called ''epigeous'' if it grows above ground, as with the morels, while underground ascocarps, such as truffles, are termed ''hypogeous''. The structure enclosing the hymenium is divided into the types described below (apothecium, cleistothecium, etc.) and this character ''is'' important for the taxonomic classification of the fungus. Apothecia can be relatively large and fleshy, whereas the others are microscopic—about the size of flecks of ...
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Soralia
Soredia are common reproductive structures of lichens. Lichens reproduce asexually by employing simple fragmentation and production of soredia and isidia. Soredia are powdery propagules composed of fungal hyphae wrapped around cyanobacteria or green algae. These can be either scattered diffusely across the surface of the lichen's thallus Thallus (plural: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. Many of these organisms ..., or produced in localized structures called soralia. Fungal hyphae make up the basic body structure of lichen. The soredia are released through openings in the upper cortex of the lichen structure. After their release, the soredia disperse to establish the lichen in a new location. References Fungal morphology and anatomy Lichenology {{lichen-stub ...
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Medulla (lichenology)
The medulla is a horizontal layer within a lichen thallus. It is a loosely arranged layer of interlaced hyphae below the upper cortex and photobiont A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Galloway, D.J. (1992). Flora of Australia - ''Lichen Glossary'' The medulla generally has a cottony appearance. It is the widest layer of a heteromerous lichen thallus.


References

Fungal morphology and anatomy Lichenology {{lichen-stub ...
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Limestone
Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms when these minerals precipitate out of water containing dissolved calcium. This can take place through both biological and nonbiological processes, though biological processes, such as the accumulation of corals and shells in the sea, have likely been more important for the last 540 million years. Limestone often contains fossils which provide scientists with information on ancient environments and on the evolution of life. About 20% to 25% of sedimentary rock is carbonate rock, and most of this is limestone. The remaining carbonate rock is mostly dolomite, a closely related rock, which contains a high percentage of the mineral dolomite, . ''Magnesian limestone'' is an obsolete and poorly-defined term used variously for dolomite, for limes ...
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Crustose Lichen
Crustose lichens are lichens that form a crust which strongly adheres to the Substrate (biology), substrate (soil, rock, tree bark, etc.), making separation from the substrate impossible without destruction. The basic structure of crustose lichens consists of a cortex (botany), cortex layer, an algal layer, and a medulla. The upper cortex layer is differentiated and is usually pigmented. The algal layer lies beneath the cortex. The medulla fastens the lichen to the substrate and is made up of Fungus, fungal hyphae. The surface of crustose lichens is characterized by branching cracks that periodically close in response to climatic variations such as alternate wetting and drying regimes. Subtypes * Powdery – considered as the simplest subtype due to the absence of an organized thallus. :The thallus appears powdery. :E.g. Genera ''Lepraria'', ''Vezdaea'' * Endolithic – grows inside the rock, usually in interstitial spaces between mineral grains. The :upper cortex is usually d ...
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Eocene
The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "dawn") and (''kainós'', "new") and refers to the "dawn" of modern ('new') fauna that appeared during the epoch. The Eocene spans the time from the end of the Paleocene Epoch to the beginning of the Oligocene Epoch. The start of the Eocene is marked by a brief period in which the concentration of the carbon isotope Carbon-13, 13C in the atmosphere was exceptionally low in comparison with the more common isotope Carbon-12, 12C. The end is set at a major extinction event called the ''Grande Coupure'' (the "Great Break" in continuity) or the Eocene–Oligocene extinction event, which may be related to the impact of one or more large bolides in Popigai impact structure, Siberia and in what is now ...
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