Cynognathia
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Cynognathia
Cynognathia ("dog jaw") is one of two major clades of cynodonts, the other being Probainognathia. Cynognathians included the large carnivorous genus ''Cynognathus'' and the herbivorous traversodontids. Cynognathians can be identified by several synapomorphies including a very deep zygomatic arch that extends above the middle of the orbit. Cynognathian fossils are currently known from Africa, Antarctica, Asia, Europe, North America and South America. Taxonomy *Suborder Cynodontia **Infraorder Eucynodontia ***(unranked) Cynognathia ****Family Cynognathidae *****''Cynognathus'' ****(unranked) Gomphodontia *****Family Diademodontidae ******''Diademodon'' ******'' Titanogomphodon'' *****(unranked) Neogomphodontia ******Family Trirachodontidae *******Subfamily Trirachodontinae ********''Langbergia'' ********'' Trirachodon'' *******Subfamily Sinognathinae ********''Beishanodon'' ********'' Sinognathus'' ********''Cricodon'' ******Family Traversodontidae *******'' Etjoia'' *******'' ...
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Cynognathus Crateronotus
''Cynognathus'' is an extinct genus of large-bodied cynodontian therapsids that lived in the Middle Triassic. It is known from a single species, ''Cynognathus crateronotus''. ''Cynognathus'' was a long predator closely related to mammals and had a southern hemispheric distribution. Fossils have so far been recovered from South Africa, Argentina, Antarctica, and Namibia. Description ''Cynognathus'' was a heavily built animal, and measured around in snout-to-vent body length. It had a particularly large head, up to in length, with wide jaws and sharp teeth. Its hindlimbs were placed directly beneath the body, but the forelimbs sprawled outwards in a more reptilian fashion. This form of double (erect/sprawling) gait is also found in some primitive mammals alive today. Possible autapomorphies of ''C. crateronotus'' include an extremely elongated postorbital bar and sectorial postcanine teeth with two serrated cusps distal to a recurved apex. Discovery and naming During 18 ...
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Cynognathus
''Cynognathus'' is an extinct genus of large-bodied cynodontian therapsids that lived in the Middle Triassic. It is known from a single species, ''Cynognathus crateronotus''. ''Cynognathus'' was a long predator closely related to mammals and had a southern hemispheric distribution. Fossils have so far been recovered from South Africa, Argentina, Antarctica, and Namibia. Description ''Cynognathus'' was a heavily built animal, and measured around in snout-to-vent body length. It had a particularly large head, up to in length, with wide jaws and sharp teeth. Its hindlimbs were placed directly beneath the body, but the forelimbs sprawled outwards in a more reptilian fashion. This form of double (erect/sprawling) gait is also found in some primitive mammals alive today. Possible autapomorphies of ''C. crateronotus'' include an extremely elongated postorbital bar and sectorial postcanine teeth with two serrated cusps distal to a recurved apex. Discovery and naming During 18 ...
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Cynodont
The cynodonts () (clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variety of lifestyles, including carnivory and herbivory. Mammals are cynodonts, as are their extinct ancestors and close relatives, having evolved from advanced probainognathian cynodonts during the Late Triassic. All other cynodont lines went extinct, with the last known non-mammalian cynodont group, the Tritylodontidae, having its youngest records in the Early Cretaceous. Description Early cynodonts have many of the skeletal characteristics of mammals. The teeth were fully differentiated and the braincase bulged at the back of the head. Outside of some crown-group mammals (notably the therians), all cynodonts probably laid eggs. The temporal fenestrae were much larger than those of their ancestors, and the widening of the zygomatic arch in ...
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Trirachodontinae
Trirachodontidae is an extinct, possibly paraphyletic family of cynognathian cynodonts from the Triassic of China and southern Africa. Trirachodontids appeared during the Early Triassic soon after the Permian-Triassic extinction event and quickly spread over a wide geographic area in a comparatively brief amount of time from 250 to 237 million years ago. Trirachodontids have wide skulls and short, narrow snouts. Two large holes called temporal fenestrae run along the back of the head and have a uniformly large width. Trirachodontids also have two large canine teeth and smaller cusped postcanines. Most of the features that distinguish trirachodonts from other cynodonts are found in their dentition. Trirachodontids lived in semi-arid environments with seasonal rainfall. The bone structure of trirachodontids suggests that they grew quickly in seasons with high rainfall and slowly in less favorable seasons. One trirachodontid, '' Trirachodon'', has been found in association with com ...
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Gomphodontia
Gomphodontia is a clade of cynognathian cynodonts that includes the families Diademodontidae, Trirachodontidae, and Traversodontidae. Gomphodonts are distinguished by wide and closely spaced molar-like postcanine teeth, which are convergent with those of mammals. Other distinguishing characteristics of gomphodonts include deep zygomatic arches, upper postcanines with three or more cusps spanning their widths and lower postcanines with two cusps spanning their widths. Gomphodonts first appeared in the Early Triassic and became extinct at the end of the Late Triassic. Fossils are known from southern Africa, Argentina and southern Brazil (Paleorrota geopark), eastern North America, Europe, China, and Antarctica. Gomphodontia was first named by paleontologist Harry Seeley in 1895. He considered it an order of wide-toothed therapsids (then called anomodonts) from South Africa, distinct from Cynodontia. By the 1930s Gomphodontia was considered a suborder of Cynodontia and included the ...
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Trirachodontidae
Trirachodontidae is an extinct, possibly paraphyletic family of cynognathian cynodonts from the Triassic of China and southern Africa. Trirachodontids appeared during the Early Triassic soon after the Permian-Triassic extinction event and quickly spread over a wide geographic area in a comparatively brief amount of time from 250 to 237 million years ago. Trirachodontids have wide skulls and short, narrow snouts. Two large holes called temporal fenestrae run along the back of the head and have a uniformly large width. Trirachodontids also have two large canine teeth and smaller cusped postcanines. Most of the features that distinguish trirachodonts from other cynodonts are found in their dentition. Trirachodontids lived in semi-arid environments with seasonal rainfall. The bone structure of trirachodontids suggests that they grew quickly in seasons with high rainfall and slowly in less favorable seasons. One trirachodontid, '' Trirachodon'', has been found in association with com ...
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Sinognathinae
Trirachodontidae is an extinct, possibly paraphyletic family of cynognathian cynodonts from the Triassic of China and southern Africa. Trirachodontids appeared during the Early Triassic soon after the Permian-Triassic extinction event and quickly spread over a wide geographic area in a comparatively brief amount of time from 250 to 237 million years ago. Trirachodontids have wide skulls and short, narrow snouts. Two large holes called temporal fenestrae run along the back of the head and have a uniformly large width. Trirachodontids also have two large canine teeth and smaller cusped postcanines. Most of the features that distinguish trirachodonts from other cynodonts are found in their dentition. Trirachodontids lived in semi-arid environments with seasonal rainfall. The bone structure of trirachodontids suggests that they grew quickly in seasons with high rainfall and slowly in less favorable seasons. One trirachodontid, '' Trirachodon'', has been found in association with com ...
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Neogomphodontia
Gomphodontia is a clade of cynognathian cynodonts that includes the families Diademodontidae, Trirachodontidae, and Traversodontidae. Gomphodonts are distinguished by wide and closely spaced molar-like postcanine teeth, which are convergent with those of mammals. Other distinguishing characteristics of gomphodonts include deep zygomatic arches, upper postcanines with three or more cusps spanning their widths and lower postcanines with two cusps spanning their widths. Gomphodonts first appeared in the Early Triassic and became extinct at the end of the Late Triassic. Fossils are known from southern Africa, Argentina and southern Brazil (Paleorrota geopark), eastern North America, Europe, China, and Antarctica. Gomphodontia was first named by paleontologist Harry Seeley in 1895. He considered it an order of wide-toothed therapsids (then called anomodonts) from South Africa, distinct from Cynodontia. By the 1930s Gomphodontia was considered a suborder of Cynodontia and included the ...
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Traversodontid
Traversodontidae is an extinct family of herbivorous cynodonts. Traversodonts were primarily Gondwanan, with many species known from Africa and South America. Recently, traversodonts have also been found from Europe and eastern North America. Traversodonts first appeared in the Middle Triassic and diversified in the Late Triassic before going extinct at the end of the epoch. The family Traversodontidae was erected by Friedrich von Huene in 1936 for cynodonts first found in São Pedro do Sul in Paleorrota, Brazil. Description Traversodonts are members of Gomphodontia, a group of herbivorous cynognathian cynodonts. As an adaptation toward eating plants, they have wide postcanine teeth behind large canines. These postcanines are closely spaced with their crowns touching each other. Each is usually wider than it is long and is covered in several cusps. Because of their complexity, postcanine teeth are the primary means of identifying and distinguishing different species of traverso ...
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Traversodontidae
Traversodontidae is an extinct family of herbivorous cynodonts. Traversodonts were primarily Gondwanan, with many species known from Africa and South America. Recently, traversodonts have also been found from Europe and eastern North America. Traversodonts first appeared in the Middle Triassic and diversified in the Late Triassic before going extinct at the end of the epoch. The family Traversodontidae was erected by Friedrich von Huene in 1936 for cynodonts first found in São Pedro do Sul in Paleorrota, Brazil. Description Traversodonts are members of Gomphodontia, a group of herbivorous cynognathian cynodonts. As an adaptation toward eating plants, they have wide postcanine teeth behind large canines. These postcanines are closely spaced with their crowns touching each other. Each is usually wider than it is long and is covered in several cusps. Because of their complexity, postcanine teeth are the primary means of identifying and distinguishing different species of traverso ...
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Probainognathia
Probainognathia is one of the two major subgroups of the clade Eucynodontia, the other being Cynognathia. The earliest forms were carnivorous and insectivorous, though some groups eventually also evolved herbivorous diets. The earliest and most basal probainognathian is the Middle Triassic (Anisian) aged ''Lumkuia'', from South Africa, though probainognathians would not become prominent until the mid Norian stage of the Late Triassic. Three groups survived the extinction at the end of Triassic: Tritheledontidae and Tritylodontidae, which both survived until the Jurassic—the latter even into the Cretaceous (''Montirictus'' and '' Xenocretosuchus'')—and Mammaliaformes, which includes the mammals. Phylogeny Below is a cladogram from Ruta, Botha-Brink, Mitchell and Benton (2013) showing one hypothesis of cynodont relationships: See also * Evolution of mammals * List of prehistoric mammals This is an incomplete list of prehistoric mammals. It does not include extant mam ...
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Eucynodontia
Eucynodontia ("true dog teeth") is a clade of cynodont therapsids including mammals and most non-mammalian cynodonts. The oldest eucynodonts are known from the Early Triassic and possibly Late Permian. Eucynodontia includes two major subgroups, Cynognathia and Probainognathia. The clade was named in 1982 by Thomas Kemp, who defined it as all cynodonts more derived than ''Thrinaxodon''. In 2001, Hopson and Kitching redefined the clade Eucynodontia as the least inclusive group containing Mammalia and ''Exaeretodon ''Exaeretodon'' is an extinct genus of fairly large, low-slung traversodontid cynodonts from the southern parts of Pangea. Four species are known, from various formations. ''E. argentinus'' is from the Carnian-age (Late Triassic) Cancha de Boch ...''.James A. Hopson and James W. Kitching, 2001, "A Probainognathian Cynodont from South Africa and the Phylogeny of Nonmammalian Cynodonts" pp 5-35 in: PARISH A. JENKINS, JR., MICHAEL D. SHAPIRO, AND TOMASZ OWERKOWICZ, ...
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