Coptoclavidae
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Coptoclavidae
Coptoclavidae is an extinct family of aquatic beetles in the suborder Adephaga. The Coptoclavidae lived from the Late Triassic to the Early Cretaceous. Coptoclavidae is a member of the adephagan clade Dytiscoidea, which contains other aquatic beetles. Suggested reasons for their extinction to include the rise of teleost fish, or competition with Gyrinidae and Dytiscidae, which possess defensive secretions and sucking channels in the mandibles of larvae, which coptoclavids likely lacked. It has been suggested that the genus '' Timarchopsis'' and the subfamily Timarchopsinae are only distantly related to other coptoclavids based on cladistic analysis, with ''Timarchopsis'' being more closely related to geadephagans like carabids and trachypachids instead. Another study also suggested similarly for '' Coptoclavisca'' and possibly other ''c''optoclaviscines. Taxonomy * †'' Agrascapha'' Lin 1992 Huangshanjie Formation, China, Carnian * †'' Amblycephalonius'' Bode 1953 Posid ...
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Coptoclavinae
Coptoclavinae is a beetle subfamily in the family Coptoclavidae Coptoclavidae is an extinct family of aquatic beetles in the suborder Adephaga. The Coptoclavidae lived from the Late Triassic to the Early Cretaceous. Coptoclavidae is a member of the adephagan clade Dytiscoidea, which contains other aquatic b .... References External links * Coptoclavidae {{Adephaga-stub ...
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Hispanoclavininae
Hispanoclavinae is an extinct subfamily of ground and water beetles in the family Coptoclavidae Coptoclavidae is an extinct family of aquatic beetles in the suborder Adephaga. The Coptoclavidae lived from the Late Triassic to the Early Cretaceous. Coptoclavidae is a member of the adephagan clade Dytiscoidea, which contains other aquatic b .... References External links * Coptoclavidae {{Adephaga-stub ...
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Adephaga
The Adephaga (from Greek ἀδηφάγος, ''adephagos'', "gluttonous") are a suborder of beetles, and with more than 40,000 recorded species in 10 families, the second-largest of the four beetle suborders. Members of this suborder are collectively known as adephagans. The largest family is Carabidae (ground beetles) which comprises most of the suborder with over 40,000 species. Adephaga also includes a variety of aquatic beetles, such as predaceous diving beetles and whirligig beetles. Anatomy Adephagans have simple antennae with no pectination or clubs. The galeae of the maxillae usually consist of two segments. Adult adephagans have visible notopleural sutures. The first visible abdominal sternum is completely separated by the hind coxae, which is one of the most easily recognizable traits of adephagans. Five segments are on each foot. Wings The transverse fold of the hind wing is near the wing tip. The median nervure ends at this fold, where it is joined by a cro ...
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Beetle
Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 described species, is the largest of all orders, constituting almost 40% of described insects and 25% of all known animal life-forms; new species are discovered frequently, with estimates suggesting that there are between 0.9 and 2.1 million total species. Found in almost every habitat except the sea and the polar regions, they interact with their ecosystems in several ways: beetles often feed on plants and fungi, break down animal and plant debris, and eat other invertebrates. Some species are serious agricultural pests, such as the Colorado potato beetle, while others such as Coccinellidae (ladybirds or ladybugs) eat aphids, scale insects, thrips, and other plant-sucking insects that damage crops. Beetles typically have a particularly hard e ...
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Trachypachidae
The Trachypachidae (sometimes known as false ground beetles) are a family of beetles that generally resemble small ground beetles, but that are distinguished by the large coxae of their rearmost legs. There are only six known extant species in the family, with four species of ''Trachypachus'' found in northern Eurasia and northern North America, and two species of ''Systolosoma'' in Chile and Argentina. They were much more diverse in the past, with dozens of described species from the Mesozoic. Their habits are similar to those of the ground beetles; they are usually found in the leaf litter of conifer forests, and are thought to be mostly predators and scavengers, active during the day ( diurnal). According to entomologist George Ball, "the most interesting thing about this small family is its uncertain phylogenetic relationships"; it has previously been classified as a subfamily of Carabidae, and as a family been placed in several different positions relative to other beetle ...
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Carnian
The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followed by the Norian. Its boundaries are not characterized by major extinctions or biotic turnovers, but a climatic event (known as the Carnian pluvial episode characterized by substantial rainfall) occurred during the Carnian and seems to be associated with important extinctions or biotic radiations. Stratigraphic definitions The Carnian was named in 1869 by Mojsisovics. It is unclear if it was named after the Carnic Alps or after the Austrian region of Carinthia (''Kärnten'' in German) or after the Carnia historical region in northwestern Italy. The name, however, was first used referring to a part of the Hallstatt Limestone cropping out in Austria. The base of the Carnian Stage is defined as the place in the stratigraphic record where t ...
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Cladistic Analysis
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived characteristics (synapomorphies'')'' that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms ''worms'' or ''fishes'' were used within a ''strict'' cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically, outside of cladistics, e.g. as a 'grade', which are fruitless to precisely delineate, especially when including extinct species. Radi ...
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