Chordate
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Chordate
A chordate () is an animal of the phylum Chordata (). All chordates possess, at some point during their larval or adult stages, five synapomorphies, or primary physical characteristics, that distinguish them from all the other taxa. These five synapomorphies include a notochord, dorsal hollow nerve cord, endostyle or thyroid, pharyngeal slits, and a post-anal tail. The name “chordate” comes from the first of these synapomorphies, the notochord, which plays a significant role in chordate structure and movement. Chordates are also Bilateral symmetry, bilaterally symmetric, have a coelom, possess a circulatory system, and exhibit Metameric, metameric segmentation. In addition to the morphological characteristics used to define chordates, analysis of genome sequences has identified two conserved signature indels (CSIs) in their proteins: cyclophilin-like protein and mitochondrial inner membrane protease ATP23, which are exclusively shared by all vertebrates, tunicates and cep ...
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Chordate
A chordate () is an animal of the phylum Chordata (). All chordates possess, at some point during their larval or adult stages, five synapomorphies, or primary physical characteristics, that distinguish them from all the other taxa. These five synapomorphies include a notochord, dorsal hollow nerve cord, endostyle or thyroid, pharyngeal slits, and a post-anal tail. The name “chordate” comes from the first of these synapomorphies, the notochord, which plays a significant role in chordate structure and movement. Chordates are also Bilateral symmetry, bilaterally symmetric, have a coelom, possess a circulatory system, and exhibit Metameric, metameric segmentation. In addition to the morphological characteristics used to define chordates, analysis of genome sequences has identified two conserved signature indels (CSIs) in their proteins: cyclophilin-like protein and mitochondrial inner membrane protease ATP23, which are exclusively shared by all vertebrates, tunicates and cep ...
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Pikaiidae
''Pikaia gracilens'' is an extinct, primitive chordate animal known from the Middle Cambrian Burgess Shale of British Columbia. Described in 1911 by Charles Doolittle Walcott as an annelid, and in 1979 by Harry B. Whittington and Simon Conway Morris as a chordate, it became the "most famous early chordate fossil," or "famously known as the earliest" chordates. It is estimated to have lived during the latter period of the Cambrian explosion. Since it initial discovery, more than a hundred specimens have been recovered. The body structure resembles that of the lancelet and perhaps it swam much like an eel. A notochord and myomeres (segmented blocks of skeletal muscles) span the entire length of the body, and are considered as the defining signatures of chordate characters. Its primitive feature is indicated by the body covering, a cuticle, which is characteristic of invertebrates and some protochordates. The exact phylogenetic position is unclear. Proposed affinities include th ...
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Metaspriggiidae
''Metaspriggina'' is a genus of chordate initially known from two specimens in the Middle Cambrian Burgess Shale and 44 specimens found in 2012 at the Marble Canyon bed in Kootenay National Park. Whilst named after the Ediacaran organism ''Spriggina'', later work has shown the two to be unrelated. ''Metaspriggina'' is considered to represent a primitive chordate, possibly transitional between cephalochordates and the earliest vertebrates, albeit this has been questioned because it seems to possess most of the characteristics attributed to craniates. It lacked fins and it had a weakly developed cranium, but it did possess two well-developed upward-facing eyes with nostrils behind them. ''Metaspriggina'' also possessed a notochord, along with seven pairs of pharyngeal bars, possibly made of cartilage. Surprisingly they were not formed from a singular bone, but they were formed of multiple separate pairs of bones, along with first two of them that were enlarged compared to th ...
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Endostyle
The endostyle is an anatomical feature found in invertebrate chordates and larval lampreys. It is an organ which assists chordates in filter-feeding. It is found in adult urochordates and cephalochordates, as well as in the larvae of the vertebrate lampreys, but has been transformed through evolutionary time into the thyroid in all other vertebrates. Since the endostyle is found in all three branches of chordates, it is presumed to have arisen in the common ancestor of these taxa, along with a shift to internal feeding for extracting suspended food from the water. The endostyle is found in the pharynx. The food particles suspended in the water adhere to the mucus it produces. The filtered water is expelled through the gill slits, while the food and mucus is then passed, by the sweeping movement of cilia that coats the endostyle, through the pharynx of the organism and into the esophagus. The endostyle in larval lampreys (ammocoetes) metamorphoses into the thyroid gland in adults ...
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Animal
Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motility, able to move, can Sexual reproduction, reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of Cell (biology), cells, the blastula, during Embryogenesis, embryonic development. Over 1.5 million Extant taxon, living animal species have been Species description, described—of which around 1 million are Insecta, insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have Ecology, complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a Symmetry in biology#Bilate ...
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Cephalochordate
A cephalochordate (from Greek: κεφαλή ''kephalé'', "head" and χορδή ''khordé'', "chord") is an animal in the chordate subphylum, Cephalochordata. They are commonly called lancelets. Cephalochordates possess 5 synapomorphies, or primary characteristics, that all chordates have at some point during their larval or adulthood stages. These 5 synapomorphies include a notochord, dorsal hollow nerve cord, endostyle, pharyngeal slits, and a post-anal tail (see chordate for descriptions of each). The fine structure of the cephalochordate notochord is best known for the Bahamas lancelet, ''Asymmetron lucayanum''. Cephalochordates are represented in modern oceans by the Amphioxiformes and are commonly found in warm temperate and tropical seas worldwide. With the presence of a notochord, adult amphioxus are able to swim and tolerate the tides of coastal environments, but they are most likely to be found within the sediment of these communities. Cephalochordates are segmented mar ...
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Olfactores
Olfactores is a clade within the Chordata that comprises the Tunicata (Urochordata) and the Vertebrata (sometimes referred to as Craniata). Olfactores represent the overwhelming majority of the phylum Chordata, as the Cephalochordata are the only chordates not included in the clade. This clade is defined by a more advanced olfactory system which, in the immediate vertebrate generation, caused the appearance of nostrils. A rudimentary neural crest is present in tunicates, implying its presence in the olfactores ancestor also, as vertebrates have a true neural crest. For this reason, they are also known as Cristozoa. Olfactores hypothesis While the hypothesis that Cephalochordata is a sister taxon to Craniata is of long standing and was once widely accepted—likely influenced by significant tunicate morphological apomorphies from other chordates, with cephalochordates even being nicknamed ‘honorary vertebrates’—studies since 2006 analyzing large sequencing datasets strong ...
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Vertebrate
Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, with currently about 69,963 species described. Vertebrates comprise such groups as the following: * jawless fish, which include hagfish and lampreys * jawed vertebrates, which include: ** cartilaginous fish (sharks, rays, and ratfish) ** bony vertebrates, which include: *** ray-fins (the majority of living bony fish) *** lobe-fins, which include: **** coelacanths and lungfish **** tetrapods (limbed vertebrates) Extant vertebrates range in size from the frog species ''Paedophryne amauensis'', at as little as , to the blue whale, at up to . Vertebrates make up less than five percent of all described animal species; the rest are invertebrates, which lack vertebral columns. The vertebrates traditionally include the hagfish, which do no ...
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Dorsal Nerve Cord
The dorsal nerve cord is a unique feature to chordates, and it is mainly found in the Vertebrata chordate subphylum. The dorsal nerve cord is only one embryonic feature unique to all chordates, among the other four chordate features-- a notochord, a post-anal tail, an endostyle, and pharyngeal slits. The dorsal hollow nerve cord is a hollow cord dorsal to the notochord. It is formed from a part of the ectoderm that rolls, forming the hollow tube. This is important, as it distinguishes chordates from other animal phyla, such as Annelids and Arthropods, which have solid, ventral tubes. The process by which this is performed is called invagination. The cells essentially convolute into the body cavity, arranging themselves on the dorsal plane above the notochord, as mentioned above. The evolutionary explanation to this adaptation from a solid cord to hollow tube is unknown. In vertebrates, the dorsal nerve cord is modified into the central nervous system, which comprises the brain and s ...
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Notochord
In anatomy, the notochord is a flexible rod which is similar in structure to the stiffer cartilage. If a species has a notochord at any stage of its life cycle (along with 4 other features), it is, by definition, a chordate. The notochord consists of inner, vacuolated cells covered by fibrous and elastic sheaths, lies along the anteroposterior axis (''front to back''), is usually closer to the dorsal than the ventral surface of the embryo, and is composed of cells derived from the mesoderm. The most commonly cited functions of the notochord are: as a midline tissue that provides directional signals to surrounding tissue during development, as a skeletal (structural) element, and as a vertebral precursor. In lancelets the notochord persists throughout life as the main structural support of the body. In tunicates the notochord is present only in the larval stage, being completely absent in the adult animal. In these invertebrate chordates, the notochord is not vacuolated. In all ...
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Phylum
In biology, a phylum (; plural: phyla) is a level of classification or taxonomic rank below kingdom and above class. Traditionally, in botany the term division has been used instead of phylum, although the International Code of Nomenclature for algae, fungi, and plants accepts the terms as equivalent. Depending on definitions, the animal kingdom Animalia contains about 31 phyla, the plant kingdom Plantae contains about 14 phyla, and the fungus kingdom Fungi contains about 8 phyla. Current research in phylogenetics is uncovering the relationships between phyla, which are contained in larger clades, like Ecdysozoa and Embryophyta. General description The term phylum was coined in 1866 by Ernst Haeckel from the Greek (, "race, stock"), related to (, "tribe, clan"). Haeckel noted that species constantly evolved into new species that seemed to retain few consistent features among themselves and therefore few features that distinguished them as a group ("a self-contained unity" ...
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Zhongxiniscus
''Zhongxiniscus'' is a genus of primitive chordate from eastern Yunnan that lived during the Early Cambrian. Known from a single specimen, it had a small, broad and short, fish-like body that was roughly ten millimeters in length. It possessed S-shaped myomeres, numbering roughly seven per one millimeter of length. Two triangular fins are evident on the dorsal margin. Evolutionary relationships When compared to other Cambrian chordates like the possible cephalochordate ''Cathaymyrus'' and the two known vertebrates ''Haikouichthys'' and ''Myllokunmingia'', ''Zhongxiniscus'' resembles ''Cathaymyrus'' in having S-shaped myomeres, but is different in having a dorsal fin and a shorter body. ''Zhongxiniscus'' approaches in form to ''Haikouichthys'' and ''Myllokunmingia'' in its myomeres and its dorsal fin, but differs from the latter two having zigzag myomeres and fin rays, which ''Zhongxiniscus'' evidently lacks. For these reasons, ''Zhongxiniscus'' is tentatively considered to be an ...
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