Charophyte
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Charophyte
Charophyta () is a group of freshwater green algae, called charophytes (), sometimes treated as a phylum, division, yet also as a superdivision or an unranked clade. The terrestrial plants, the Embryophyta emerged within Charophyta, possibly from terrestrial unicellular charophytes, with the class Zygnematophyceae as a sister group. The clade Streptophyta may be formed by placing Embryophyta within the Charophyta. The Embryophyta may already be included in the Charophyta, in which case it is a synonym of the Streptophyta. The sister group of the charophytes are the Chlorophyta. In some charophyte groups, such as the Zygnematophyceae or conjugating green algae, Flagellum, flagella are absent and sexual reproduction does not involve free-swimming flagellate sperm. Flagellate sperm, however, are found in stoneworts (Charales) and Coleochaetales, orders of parenchymatous charophytes that are the closest relatives of the land plants, where flagellate sperm are also present in all exce ...
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Green Algae
The green algae (singular: green alga) are a group consisting of the Prasinodermophyta and its unnamed sister which contains the Chlorophyta and Charophyta/Streptophyta. The land plants (Embryophytes) have emerged deep in the Charophyte alga as sister of the Zygnematophyceae. Since the realization that the Embryophytes emerged within the green algae, some authors are starting to properly include them. The completed clade that includes both green algae and embryophytes is monophyletic and is referred to as the clade Viridiplantae and as the kingdom Plantae. The green algae include unicellular and colonial flagellates, most with two flagella per cell, as well as various colonial, coccoid and filamentous forms, and macroscopic, multicellular seaweeds. There are about 22,000 species of green algae. Many species live most of their lives as single cells, while other species form coenobia (colonies), long filaments, or highly differentiated macroscopic seaweeds. A few other organi ...
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Stonewort
Charales is an order of freshwater green algae in the division Charophyta, class Charophyceae, commonly known as stoneworts. Depending on the treatment of the genus ''Nitellopsis'', living (extant) species are placed into either one family ( Characeae) or two (Characeae and Feistiellaceae). Further families are used for fossil members of the order. Linnaeus established the genus ''Chara (alga), Chara'' in 1753. Taxonomy The higher level classification of green algae was unsettled . AlgaeBase places Charales within the class Charophyceae and its Circumscription (taxonomy), circumscription of the Phylum, division Charophyta. Families The number of families and their division into genera varies. , AlgaeBase accepts two families containing some extant species and four families containing only fossil species: * Characeae S.F.Gray * Feistiellaceae Schudack *†Aclistocharaceae X.G.Zhou (may be included in Characeae) *†Atopocharaceae R.E.Peck (may be included in Clavatoraceae) *â ...
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Octochara
''Octochara'' is a genus of fossil charophyte (aquatic green alga) from the Famennian (Late Devonian). It is one of two genera of charophyte described from the Waterloo Farm lagerstätte in southern Africa. It and ''Hexachara'', from the same locality provide the oldest record of reconstruct able charophytes with in situ oogonia. ''Octochara'' is derived from a Greek word "octo", meaning eight, a reference to the octoradial symmetry, and "chara", referring to membership of the Charales. In ''Octochara'', a whorl of eight laterals are borne at each node. Each lateral is branched to produce four secondary branches and bears an oogonium An oogonium (plural oogonia) is a small diploid cell which, upon maturation, forms a primordial follicle in a female fetus or the female (haploid or diploid) gametangium of certain thallophytes. In the mammalian fetus Oogonia are formed in larg .... Two species of ''Octochara'' have been described, ''Octochara crassa'' and ''Octochara gracilis''. ...
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Charales
Charales is an order of freshwater green algae in the division Charophyta, class Charophyceae, commonly known as stoneworts. Depending on the treatment of the genus ''Nitellopsis'', living (extant) species are placed into either one family ( Characeae) or two (Characeae and Feistiellaceae). Further families are used for fossil members of the order. Linnaeus established the genus '' Chara'' in 1753. Taxonomy The higher level classification of green algae was unsettled . AlgaeBase places Charales within the class Charophyceae and its circumscription of the division Charophyta. Families The number of families and their division into genera varies. , AlgaeBase accepts two families containing some extant species and four families containing only fossil species: * Characeae S.F.Gray * Feistiellaceae Schudack *† Aclistocharaceae X.G.Zhou (may be included in Characeae) *† Atopocharaceae R.E.Peck (may be included in Clavatoraceae) *† Clavatoraceae Pia *† Porocharaceae Grambas ...
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Phylum
In biology, a phylum (; plural: phyla) is a level of classification or taxonomic rank below kingdom and above class. Traditionally, in botany the term division has been used instead of phylum, although the International Code of Nomenclature for algae, fungi, and plants accepts the terms as equivalent. Depending on definitions, the animal kingdom Animalia contains about 31 phyla, the plant kingdom Plantae contains about 14 phyla, and the fungus kingdom Fungi contains about 8 phyla. Current research in phylogenetics is uncovering the relationships between phyla, which are contained in larger clades, like Ecdysozoa and Embryophyta. General description The term phylum was coined in 1866 by Ernst Haeckel from the Greek (, "race, stock"), related to (, "tribe, clan"). Haeckel noted that species constantly evolved into new species that seemed to retain few consistent features among themselves and therefore few features that distinguished them as a group ("a self-contained unity" ...
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Rhynie Chert
The Rhynie chert is a Lower Devonian sedimentary deposit exhibiting extraordinary fossil detail or completeness (a Lagerstätte). It is exposed near the village of Rhynie, Aberdeenshire, Scotland; a second unit, the Windyfield chert, is located some 700 m away. The Rhynie chert contains exceptionally preserved plant, fungus, lichen and animal material preserved in place by an overlying volcanic deposit. The bulk of the Devonian fossil bed consists of primitive plants (which had water-conducting cells and sporangia, but no true leaves), along with arthropods, lichens, algae and fungi. This fossil bed is remarkable for two reasons. First, the age of the site ( Pragian, Early Devonian, formed about ) places it at an early stage in the colonisation of land. Second, these cherts are famous for their exceptional state of ultrastructural preservation, with individual cell walls easily visible in polished specimens. Stomata have been counted and lignin remnants detected in the ...
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Phragmoplastophyta
The Phragmoplastophyta (Lecointre & Guyader 2006) are a proposed sister clade of the Klebsormidiaceae in the Streptophyte/Charophyte clade. The Phragmoplastophyta consist of the Charophycaea and another unnamed clade which contains the Coleochaetophyceae, Zygnematophyceae, Mesotaeniaceae, and Embryophytes (land plants). It is an important step in the emergence of land plants within the green algae. It is equivalent to the ZCC clade/grade, cladistically granting the Embryophyta. The mitosis of Phragmoplastophyta takes place via a phragmoplast. Another synapomorphy In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to hav ... of this clade is the synthesis of cellulose microfibrils by a complex of octameric cellulose synthetases. This complex crosses the plasma membrane and polymerizes mole ...
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Chlorokybophyceae
''Chlorokybus'' is a multicellular (sarcinoid) genus of basal green algae or charophyte, a soil alga Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mi .... It has been classified as the sole member of the family Chlorokybaceae, which is the sole member of the order Chlorokybales, in turn the sole member of the class Chlorokybophyceae. Taxonomy ''Chlorokybus atmophyticus'' was once thought to be the only species in the genus. In 2021, a study showed that there were at least four other species, morphologically indistinguishable, but with deep genomic differences, suggesting divergences possibly about 76 million years ago. ''Chlorokybus'' has been found in Eurasia, Central and South America. ''Chlorokybus'' was placed in a new class, order and family. The new class Chlorokybophyceae was ...
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Waterloo Farm Lagerstätte
The Waterloo Farm lagerstätte is a Famennian lagerstätte in South Africa that constitutes the only known record of a near-polar Devonian coastal ecosystem. History and discovery The Waterloo Farm Lagerstätte is an approximately 360 million year old Famennian (latest Devonian) fossil-rich locality of the Witpoort Formation (Witteberg Group, Cape Supergroup) in Makhanda (former Grahamstown) within the Eastern Cape Province, South Africa. Before it’s discovery very little was known of life during the Famennian (see Late Devonian extinction) in what is now southern Africa . This is largely due to the fact that fossils in the Witpoort Formation generally occur in black anaerobically deposited metashale that rapidly degrades near surface and is therefore rarely seen in natural outcrop. As is the case with many other scientific discoveries, the discovery of Waterloo Farm was accidental. Uprisings against the apartheid system in South Africa had by the mid-1980s escalated to vi ...
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Flagellum
A flagellum (; ) is a hairlike appendage that protrudes from certain plant and animal sperm cells, and from a wide range of microorganisms to provide motility. Many protists with flagella are termed as flagellates. A microorganism may have from one to many flagella. A gram-negative bacterium ''Helicobacter pylori'' for example uses its multiple flagella to propel itself through the mucus lining to reach the stomach epithelium, where it may cause a gastric ulcer to develop. In some bacteria the flagellum can also function as a sensory organelle, being sensitive to wetness outside the cell. Across the three domains of Bacteria, Archaea, and Eukaryota the flagellum has a different structure, protein composition, and mechanism of propulsion but shares the same function of providing motility. The Latin word means " whip" to describe its lash-like swimming motion. The flagellum in archaea is called the archaellum to note its difference from the bacterial flagellum. Eukaryotic ...
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Devonian
The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, where rocks from this period were first studied. The first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents. By the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. The arthropod groups of myriapods, arachnids and hexapods also became well-established early in this period, after starting their expansion to land at least from the Ordovician period. Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The placoderms began dominating ...
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Angiosperm
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants that produce their seeds enclosed within a fruit. They are by far the most diverse group of land plants with 64 orders, 416 families, approximately 13,000 known genera and 300,000 known species. Angiosperms were formerly called Magnoliophyta (). Like gymnosperms, angiosperms are seed-producing plants. They are distinguished from gymnosperms by characteristics including flowers, endosperm within their seeds, and the production of fruits that contain the seeds. The ancestors of flowering plants diverged from the common ancestor of all living gymnosperms before the end of the Carboniferous, over 300 million years ago. The closest fossil relatives of flowering plants are uncertain and contentious. The earliest angiosperm fossils are in the ...
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