Calciavis Grandei
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Calciavis Grandei
''Calciavis'' is an extinct genus of bird from the Eocene of Wyoming. It is a lithornithid, a member of a lineage of flying palaeognaths, distantly related to modern ratites. Like many other fossils from the Green River Formation, it is exceptionally well preserved, bearing soft-tissue impressions of skin and feathers. Discovery and Description The original specimen of ''Calciavis'', AMNH 30578, was found in the Fossil Butte Member of the Green River Formation, Wyoming, dating to around 51 million years ago. It is an exceptionally well preserved specimen, preserving most of the post-cranial skeleton aside from the femur and associated pelvic region, as well as soft-tissue impressions of feathers, skin, Bird anatomy#Scales, foot scales and claw sheaths. Another specimen, AMNH 30560 , was found in the vicinity, composed of a similarly complete most mostly disarticulate skeleton. It is diagnosed by several features of the skull and pelvis as well as having metatarsals IV and II sube ...
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Eocene
The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "dawn") and (''kainós'', "new") and refers to the "dawn" of modern ('new') fauna that appeared during the epoch. The Eocene spans the time from the end of the Paleocene Epoch to the beginning of the Oligocene Epoch. The start of the Eocene is marked by a brief period in which the concentration of the carbon isotope Carbon-13, 13C in the atmosphere was exceptionally low in comparison with the more common isotope Carbon-12, 12C. The end is set at a major extinction event called the ''Grande Coupure'' (the "Great Break" in continuity) or the Eocene–Oligocene extinction event, which may be related to the impact of one or more large bolides in Popigai impact structure, Siberia and in what is now ...
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Tarsometatarsus
The tarsometatarsus is a bone that is only found in the lower leg of birds and some non-avian dinosaurs. It is formed from the fusion of several bones found in other types of animals, and homologous to the mammalian tarsus (ankle bones) and metatarsal bones (foot). Despite this, the tarsometatarsus of birds is often referred to as just the shank, tarsus or metatarsus. Tarsometatarsal fusion occurred in several ways and extents throughout bird evolution. Specifically, in Neornithes (modern birds), although the bones are joined along their entire length, the fusion is most thorough at the distal (metatarsal) end. In the Enantiornithes, a group of Mesozoic avialans, the fusion was complete at the proximal (tarsal) end, but the distal metatarsi were still partially distinct. While these fused bones are best known from birds and their relatives, avians are neither the only group nor the first to possess tarsometatarsi. In a remarkable case of parallel evolution, they were also pres ...
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Paleogene Birds Of North America
The Paleogene ( ; also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period million years ago ( Mya) to the beginning of the Neogene Period Mya. It is the beginning of the Cenozoic Era of the present Phanerozoic Eon. The earlier term Tertiary Period was used to define the span of time now covered by the Paleogene Period and subsequent Neogene Period; despite no longer being recognised as a formal stratigraphic term, 'Tertiary' is still widely found in earth science literature and remains in informal use. Paleogene is often abbreviated "Pg" (but the United States Geological Survey uses the abbreviation PE for the Paleogene on the Survey's geologic maps). During the Paleogene, mammals diversified from relatively small, simple forms into a large group of diverse animals in the wake of the Cretaceous–Paleogene extinction event that ended the preceding ...
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Eocene Genus Extinctions
The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "dawn") and (''kainós'', "new") and refers to the "dawn" of modern ('new') fauna that appeared during the epoch. The Eocene spans the time from the end of the Paleocene Epoch to the beginning of the Oligocene Epoch. The start of the Eocene is marked by a brief period in which the concentration of the carbon isotope 13C in the atmosphere was exceptionally low in comparison with the more common isotope 12C. The end is set at a major extinction event called the ''Grande Coupure'' (the "Great Break" in continuity) or the Eocene–Oligocene extinction event, which may be related to the impact of one or more large bolides in Siberia and in what is now Chesapeake Bay. As with other geologic periods, the strata that define the start and end of the ...
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Paleogene Birds Of Europe
The Paleogene ( ; also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period million years ago ( Mya) to the beginning of the Neogene Period Mya. It is the beginning of the Cenozoic Era of the present Phanerozoic Eon. The earlier term Tertiary Period was used to define the span of time now covered by the Paleogene Period and subsequent Neogene Period; despite no longer being recognised as a formal stratigraphic term, 'Tertiary' is still widely found in earth science literature and remains in informal use. Paleogene is often abbreviated "Pg" (but the United States Geological Survey uses the abbreviation PE for the Paleogene on the Survey's geologic maps). During the Paleogene, mammals diversified from relatively small, simple forms into a large group of diverse animals in the wake of the Cretaceous–Paleogene extinction event that ended the preceding ...
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Lithornithidae
Lithornithidae is an extinct, possibly paraphyletic (but see below) group of early paleognath birds. They are known from fossils dating to the Upper Paleocene through the Middle Eocene of North America and Europe, with possible Late Cretaceous representatives. All are extinct today; the youngest specimen is the currently unnamed SGPIMH MEV1 specimen from the mid-Eocene Messel Pit site. Lithornithids had long, slender, bills for probing. They closely resembled modern tinamous. They possessed a rhynchokinetic skull with relatively unfused cranial bones, a weakly fused pygostyle and a splenial. The unguals were more curved than in tinamous and probably allowed better perching in trees. The order Lithornithiformes was erected by Dr. Peter Houde in 1988. Initially, only three genera ('' Lithornis'', '' Paracathartes'', and '' Pseudocrypturus'') and eight named species were included. '' Promusophaga'' (Harrison & Walker, 1977) originally considered a stem-turaco The turacos m ...
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Barbule
Feathers are epidermal growths that form a distinctive outer covering, or plumage, on both avian (bird) and some non-avian dinosaurs and other archosaurs. They are the most complex integumentary structures found in vertebrates and a premier example of a complex evolutionary novelty. They are among the characteristics that distinguish the extant birds from other living groups. Although feathers cover most of the bird's body, they arise only from certain well-defined tracts on the skin. They aid in flight, thermal insulation, and waterproofing. In addition, coloration helps in communication and protection. Plumology (or plumage science) is the name for the science that is associated with the study of feathers. Feathers have a number of utilitarian, cultural, and religious uses. Feathers are both soft and excellent at trapping heat; thus, they are sometimes used in high-class bedding, especially pillows, blankets, and mattresses. They are also used as filling for winter clothi ...
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Plumage
Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can be different colour morphs. The placement of feathers on a bird is not haphazard, but rather emerge in organized, overlapping rows and groups, and these are known by standardized names. Most birds moult twice a year, resulting in a breeding or ''nuptial plumage'' and a ''basic plumage''. Many ducks and some other species such as the red junglefowl have males wearing a bright nuptial plumage while breeding and a drab ''eclipse plumage'' for some months afterward. The painted bunting's juveniles have two inserted moults in their first autumn, each yielding plumage like an adult female. The first starts a few days after fledging replacing the ''juvenile plumage'' with an ''auxiliary formative plumage''; the second a month or so l ...
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Paracathartes
''Paracathartes'' is a genus of extinct bird from the Wasachtian horizon of lower Eocene Wyoming. One species, ''Paracathartes howardae'' has been described. It is a paleognathous bird, turkey-like in stature and size, that probably resembled a tinamou quite closely. ''Paracathartes'' was described by Harrison as the earliest known cathartid vulture. Rich criticized this assignment.Rich, P. V. (1983) Houde (1988) included it as a member of the order Lithornithiformes and family Lithornithidae.Houde, Peter W. (1988) The holotype specimen is in the collection of the Royal Ontario Museum. It has catalog number ROM 22658. It is the distal end of a left tibiotarsus. It was collected by G. E. Lindblad and G. Sternberg on 4 August 1949. It was found at the northernmost branch of Elk Creek, near Basin, Wyoming. The horizon is Greybullian, middle Wasatchian (early Eocene), Willwood Formation, Bighorn Basin. Further specimens of ''Paracathartes'' were collected, including almost the ...
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Ischium
The ischium () forms the lower and back region of the (''os coxae''). Situated below the ilium and behind the pubis, it is one of three regions whose fusion creates the . The superior portion of this region forms approximately one-third of the acetabulum.


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Lithornis Promiscuus
''Lithornis'' is a genus of extinct paleognathous birds. Although ''Lithornis'' was able to fly well, their closest relatives are the extant tinamous (which are poor flyers) and ratites (which are flightless birds). Fossils of ''Lithornis'' are known with certainty from the Upper Paleocene through the Middle Eocene, but their fossil record may extend to the late Cretaceous. "''Lithornis''" is from ancient Greek for "stone bird", as it is one of the first fossil birds to become widely discussed. Presumably closely related genera are ''Paracathartes'' and '' Pseudocrypturus''. Species Six species have been recognized in modern times; undescribed ones are also likely to exist. The supposed tarsometatarsus piece from which ''"Lithornis" emuinus'' was described is actually a humerus fragment of the giant pseudotooth bird ''Dasornis''.Houde, P. (1988) ''Lithornis celetius'' ''L. celetius'' is from the Bangtail Quarry, Sedan Quadrangle, Park County, Montana, USA, and was describe ...
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Coracoid
A coracoid (from Greek κόραξ, ''koraks'', raven) is a paired bone which is part of the shoulder assembly in all vertebrates except therian mammals (marsupials and placentals). In therian mammals (including humans), a coracoid process is present as part of the scapula, but this is not homologous with the coracoid bone of most other vertebrates. In other tetrapods it joins the scapula to the front end of the sternum and has a notch on the dorsal surface which, along with a similar notch on the ventral surface of the scapula, forms the socket in which the proximal end of the humerus (upper arm bone) is located. The acrocoracoid process is an expansion adjacent to this contact surface, to which the shoulderward end of the biceps brachii muscle attaches in these animals. In birds (and generally theropods and related animals), the entire unit is rigid and called scapulocoracoid. This plays a major role in bird flight. In dinosaurs the main bones of the pectoral girdle were the sca ...
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