Xenacoelomorpha
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Xenacoelomorpha
Xenacoelomorpha is a small phylum of bilaterian invertebrate animals, consisting of two sister groups: xenoturbellids and acoelomorphs. This new phylum was named in February 2011 and suggested based on morphological synapomorphies (physical appearances shared by the animals in the clade), which was then confirmed by phylogenomic analyses of molecular data (similarities in the DNA of the animals within the clade). Phylogenetics The clade (groupings of organisms based on their most recent shared/common ancestors) Xenacoelomorpha groups the Acoelomorpha and the genus ''Xenoturbella'', due to molecular studies. Initially this phylum was considered to be a member of the deuterostomes, (meaning during development, as an embryo, the anus develops first and then the mouth), but because of recent transcriptome analysis, it was concluded that phylum Xenacoelomorpha is the sister group (two closest relatives in a phylogenetic tree) to the Nephrozoa, which includes both the protostomes (w ...
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Bilaterian
The Bilateria or bilaterians are animals with bilateral symmetry as an embryo, i.e. having a left and a right side that are mirror images of each other. This also means they have a head and a tail (anterior-posterior axis) as well as a belly and a back (ventral-dorsal axis). Nearly all are bilaterally symmetrical as adults as well; the most notable exception is the echinoderms, which achieve secondary pentaradial symmetry as adults, but are bilaterally symmetrical during embryonic development. Most animals are bilaterians, excluding sponges, ctenophores, placozoans and cnidarians. For the most part, bilateral embryos are triploblastic, having three germ layers: endoderm, mesoderm, and ectoderm. Except for a few phyla (i.e. flatworms and gnathostomulids), bilaterians have complete digestive tracts with a separate mouth and anus. Some bilaterians lack body cavities (Acoelomata, acoelomates, i.e. Flatworm, Platyhelminthes, Gastrotricha and Gnathostomulida), while others display pri ...
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Deuterostomes
Deuterostomia (; in Greek) are animals typically characterized by their anus forming before their mouth during embryonic development. The group's sister clade is Protostomia, animals whose digestive tract development is more varied. Some examples of deuterostomes include vertebrates (and thus humans), sea stars, and crinoids. In deuterostomy, the developing embryo's first opening (the blastopore) becomes the anus, while the mouth is formed at a different site later on. This was initially the group's distinguishing characteristic, but deuterostomy has since been discovered among protostomes as well. This group is also known as enterocoelomates, because their coelom develops through enterocoely. The three major clades of deuterostomes are Chordata (e.g. vertebrates), Echinodermata (e.g. starfish), and Hemichordata (e.g. acorn worms). Together with Protostomia and their out-group Xenacoelomorpha, these compose the Bilateria, animals with bilateral symmetry and three germ layers. ...
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Xenoturbella
''Xenoturbella'' is a genus of very simple bilaterians up to a few centimeters long. It contains a small number of marine benthic worm-like species. The first known species (''Xenoturbella bocki'') was discovered in 1915 by Sixten Bock, but it was only properly described in 1949 by Einar Westblad. Description ''Xenoturbella'' has a very simple body plan. It consists of dorsoventrally flattened acoelomate animals, with an anterior circumferential furrow. It shows two ciliated epithelial layers: an external epidermis and an internal gastrodermis lining the simple sac-like gut. The multiciliated epiderm displays unique interconnected ciliary rootlets and mode of withdrawal and resorption of worn epidermal cells. The mouth is a mid-ventral pore leading to a gastral cavity, and there is no anus: waste is dispelled through the same opening as food is taken in. The nervous system is composed by a net of interconnected neurons beneath the epidermis, without any concentration of neuron ...
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Xenoturbellida
''Xenoturbella'' is a genus of very simple bilaterians up to a few centimeters long. It contains a small number of marine benthic worm-like species. The first known species (''Xenoturbella bocki'') was discovered in 1915 by Sixten Bock, but it was only properly described in 1949 by Einar Westblad. Description ''Xenoturbella'' has a very simple body plan. It consists of dorsoventrally flattened acoelomate animals, with an anterior circumferential furrow. It shows two ciliated epithelial layers: an external epidermis and an internal gastrodermis lining the simple sac-like gut. The multiciliated epiderm displays unique interconnected ciliary rootlets and mode of withdrawal and resorption of worn epidermal cells. The mouth is a mid-ventral pore leading to a gastral cavity, and there is no anus: waste is dispelled through the same opening as food is taken in. The nervous system is composed by a net of interconnected neurons beneath the epidermis, without any concentration of neuron ...
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Xenoturbellid
''Xenoturbella'' is a genus of very simple bilaterians up to a few centimeters long. It contains a small number of marine benthic worm-like species. The first known species (''Xenoturbella bocki'') was discovered in 1915 by Sixten Bock, but it was only properly described in 1949 by Einar Westblad. Description ''Xenoturbella'' has a very simple body plan. It consists of dorsoventrally flattened acoelomate animals, with an anterior circumferential furrow. It shows two ciliated epithelial layers: an external epidermis and an internal gastrodermis lining the simple sac-like gut. The multiciliated epiderm displays unique interconnected ciliary rootlets and mode of withdrawal and resorption of worn epidermal cells. The mouth is a mid-ventral pore leading to a gastral cavity, and there is no anus: waste is dispelled through the same opening as food is taken in. The nervous system is composed by a net of interconnected neurons beneath the epidermis, without any concentration of neuron ...
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Acoelomorpha
Acoelomorpha is a subphylum of very simple and small soft-bodied animals with planula-like features which live in marine or brackish waters. They usually live between grains of sediment, swimming as plankton, or crawling on other organisms, such as algae and corals.Cannon, L. R. G. (1986) ''Turbellaria of the World. A guide to families and genera''. Brisbane, Queensland Museum, 136 p. With the exception of two acoel freshwater species, all known Acoelomorphs are marine. Systematics Etymology The term "acoelomorph" derives from the Ancient Greek words (), the ''alpha privative'', expressing negation or absence, (), meaning "cavity", and (), meaning "form". This refers to the fact that acoelomorphs have a structure lacking a fluid-filled body cavity. Classification The subphylum Acoelomorpha is divided into two classes. There are at least 408 described species, with a majority of these falling within the Crucimusculata infraorder in Acoela. * Acoela comprise small fla ...
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Ambulacraria
Ambulacraria , or Coelomopora , is a clade of invertebrate phyla that includes echinoderms and hemichordates; a member of this group is called an ambulacrarian. Phylogenetic analysis suggests the echinoderms and hemichordates separated around 533 million years ago. The Ambulacraria are part of the deuterostomes, a larger clade that also includes the Chordata, Vetulicolia. The two living clades with representative organisms are: * Echinodermata (sea stars, sea urchins, brittle stars, sea cucumbers, feather stars, sea lilies, etc.) * Hemichordata (acorn worms, Pterobranchia, and possibly graptolites) (These together sometimes are called the ''lower deuterostomes''.) Whether the Xenacoelomorpha clade is the sister group to the Ambulacraria remains a contentious issue, with some authors arguing that the former should be placed more basally among metazoans, and other authors asserting that the best choices of phylogenetic methods support the position of Xenacoelomorpha as the sister ...
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Acoelomorph
Acoelomorpha is a subphylum of very simple and small soft-bodied animals with planula-like features which live in marine or brackish waters. They usually live between grains of sediment, swimming as plankton, or crawling on other organisms, such as algae and corals.Cannon, L. R. G. (1986) ''Turbellaria of the World. A guide to families and genera''. Brisbane, Queensland Museum, 136 p. With the exception of two acoel freshwater species, all known Acoelomorphs are marine. Systematics Etymology The term "acoelomorph" derives from the Ancient Greek words (), the ''alpha privative'', expressing negation or absence, (), meaning "cavity", and (), meaning "form". This refers to the fact that acoelomorphs have a structure lacking a fluid-filled body cavity. Classification The subphylum Acoelomorpha is divided into two classes. There are at least 408 described species, with a majority of these falling within the Crucimusculata infraorder in Acoela. * Acoela comprise small flat ...
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Nephrozoa
Nephrozoa is a major clade of bilaterians, divided into the protostomes and the deuterostomes, containing almost all animal phyla and over a million extant species. Its sister clade is the Xenacoelomorpha. The Ambulacraria (conventionally deuterostomes) was formerly thought to be sister to the Xenacoelomorpha, forming the Xenambulacraria as basal Deuterostomes, or basal Bilateria invalidating Nephrozoa and Deuterostomes in earlier studies. The coelom, the digestive tract and excretory organs (nephridia), and nerve cords developed in the Nephrozoa. It has been argued that, because protonephridia are only found in protostomes, they cannot be considered a synapomorphy of this group. This would make Nephrozoa an improper name, leaving Eubilateria as this clade's name. Chordates (which include all the vertebrates) are deuterostomes. It seems very likely that the ''Kimberella ''Kimberella'' is an extinct genus of bilaterian known only from rocks of the Ediacaran period. The s ...
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Animal
Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motility, able to move, can Sexual reproduction, reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of Cell (biology), cells, the blastula, during Embryogenesis, embryonic development. Over 1.5 million Extant taxon, living animal species have been Species description, described—of which around 1 million are Insecta, insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have Ecology, complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a Symmetry in biology#Bilate ...
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Xenoturbella Japonica
''Xenoturbella japonica'' is a marine benthic worm-like species that belongs to the genus ''Xenoturbella''. It has been discovered in western Pacific Ocean by a group of Japanese scientists from the University of Tsukuba. The species was described in 2017 in a study published in the journal BMC Evolutionary Biology, and amended in 2018. ''Xenotrubella japonica'' is known for lacking respiratory, circulatory and an excretory system. Description The etymology of the species name corresponds to the locality where the specimens were sampled. ''Xenoturbella japonica'' is in length, with a pale orange colouration. The body wall displays ring and side furrows. The mouth is orientated ventrally, just anterior to the ring furrow. The live specimen exhibits a conspicuous ventral epidermal glandular network. Tissues contain exogenous DNA corresponding to bivalve mollusks, the vesicomyid '' Acila castrensis'' and '' Nucula nucleus''. Phylogeny Comparison of mitochondrial DNA ...
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Xenambulacraria
Xenambulacraria is a proposed clade of animals with symmetry (biology)#Bilateral symmetry, bilateral symmetry as an embryo, consisting of the Xenacoelomorpha (i.e., ''Xenoturbella'' and acoelomorphs) and the Ambulacraria (i.e., echinoderms and hemichordates). If confirmed, the clade would either be the sister group to the chordates (deuterostomes monophyletic) or the sister group to all the other bilaterians (deuteristomes paraphyletic). File:Nemertodermatida species.png, Various Acoelomorpha (nemertodermatids). File:Enteropneusta.png, Various Hemichordata (enteropneusts). References

{{Animalia Controversial taxa Bilaterian taxa ...
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