Sphenacodonts
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Sphenacodonts
Sphenacodontia is a stem-based clade of derived synapsids. It was defined by Amson and Laurin (2011) as "the largest clade that includes ''Haptodus baylei'', ''Haptodus garnettensis'' and ''Sphenacodon ferox'', but not ''Edaphosaurus pogonias''". They first appear during the Late Pennsylvanian (Upper Carboniferous) epoch. From the end of the Carboniferous to the end of the Permian, most of them remained large, with only some secondarily becoming small in size. Basal Sphenacodontia constitute a transitional evolutionary series from early pelycosaurs to ancestral therapsids (which in turn were the ancestors of more advanced forms and finally the mammals). One might say that the sphenacodontians are proto-therapsids (even though there is almost a 30-million-years gap between the separation of the ancestors of therapsids from other sphenacodontians and the first appearance of therapsids in the fossil record). Characteristics The defining characteristics include a thickening o ...
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Sphenacodontoidea
Sphenacodontoidea is a node-based clade that is defined to include the most recent common ancestor of Sphenacodontidae and Therapsida and its descendants (including mammals). Sphenacodontoids are characterised by a number of synapomorphies concerninproportions of the bones of the skull and the teeth The sphenacodontoids evolved from earlier sphenacodonts such as ''Haptodus'' via a number of transitional stages of small, unspecialised pelycosaurs. Classification The following taxonomy follows Fröbisch ''et al.'' (2011) and Benson (2012) unless otherwise noted. Class Synapsida * Sphenacodontoidea ** Family †Sphenacodontidae ** Therapsida See also * Evolution of mammals References * Laurin, M. and Reisz, R. R., 1997Autapomorphies of the main clades of synapsids- Tree of Life Web Project The Tree of Life Web Project is an Internet project providing information about the diversity and phylogeny of life on Earth. This collaborative peer reviewed project began in 1995 ...
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Shashajaia
''Shashajaia'' is a genus of extinct non-mammalian synapsids from the late Carboniferous to Early Permian. It was one of the earliest members of the group, coming from the Gzhelian stage. It lived in what is now the Halgaito Formation within the larger Cutler group located in the U.S state of Utah. According to a description study, this synapsid is known from well preserved dentary and jaw fragments. ''Shashajaia'' shares many similarities to other sphenacodontids including, enlarged ( canine-like) anterior dentary teeth, a dorsoventrally deep symphysis and low-crowned, subthecodont postcanines having festooned plicidentine. The study also found that this genus is close to the evolutionary divergence of the Sphenacodontids and the Therapsids, from which mammalian synapsids (including humans) arose from. Based on studies done on its teeth, Paleontologists found that as their prey became more terrestrial, synapsids like ''Shashajaia'' adapted to life on land and grew larger teeth to ...
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Pennsylvanian (geology)
The Pennsylvanian ( , also known as Upper Carboniferous or Late Carboniferous) is, in the International Commission on Stratigraphy, ICS geologic timescale, the younger of two period (geology), subperiods (or upper of two system (stratigraphy), subsystems) of the Carboniferous Period. It lasted from roughly . As with most other geochronology, geochronologic units, the stratum, rock beds that define the Pennsylvanian are well identified, but the exact date of the start and end are uncertain by a few hundred thousand years. The Pennsylvanian is named after the U.S. state of Pennsylvania, where the coal-productive beds of this age are widespread. The division between Pennsylvanian and Mississippian (geology), Mississippian comes from North American stratigraphy. In North America, where the early Carboniferous beds are primarily marine limestones, the Pennsylvanian was in the past treated as a full-fledged geologic period between the Mississippian and the Permian. In parts of Europe, ...
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Stem-based Taxon
Phylogenetic nomenclature is a method of nomenclature for taxa in biology that uses phylogenetic definitions for taxon names as explained below. This contrasts with the traditional approach, in which taxon names are defined by a '' type'', which can be a specimen or a taxon of lower rank, and a description in words. Phylogenetic nomenclature is currently regulated by the '' International Code of Phylogenetic Nomenclature'' (''PhyloCode''). Definitions Phylogenetic nomenclature ties names to clades, groups consisting of an ancestor and all its descendants. These groups can equivalently be called monophyletic. There are slightly different ways of specifying the ancestor, which are discussed below. Once the ancestor is specified, the meaning of the name is fixed: the ancestor and all organisms which are its descendants are included in the named taxon. Listing all these organisms (i.e. providing a full circumscription) requires the full phylogenetic tree to be known. In practice, the ...
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Transitional Fossils
A transitional fossil is any fossilized remains of a life form that exhibits traits common to both an ancestral group and its derived descendant group. This is especially important where the descendant group is sharply differentiated by gross anatomy and mode of living from the ancestral group. These fossils serve as a reminder that taxonomic divisions are human constructs that have been imposed in hindsight on a continuum of variation. Because of the incompleteness of the fossil record, there is usually no way to know exactly how close a transitional fossil is to the point of divergence. Therefore, it cannot be assumed that transitional fossils are direct ancestors of more recent groups, though they are frequently used as models for such ancestors. In 1859, when Charles Darwin's ''On the Origin of Species'' was first published, the fossil record was poorly known. Darwin described the perceived lack of transitional fossils as "the most obvious and gravest objection which can be ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Permian
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the sauropsids ( reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their am ...
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Carboniferous
The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carboniferous'' means "coal-bearing", from the Latin '' carbō'' ("coal") and '' ferō'' ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern 'system' names, it was coined by geologists William Conybeare and William Phillips in 1822, based on a study of the British rock succession. The Carboniferous is often treated in North America as two geological periods, the earlier Mississippian and the later Pennsylvanian. Terrestrial animal life was well established by the Carboniferous Period. Tetrapods (four limbed vertebrates), which had originated from lobe-finned fish during the preceding Devonian, became pentadactylous in and diversified during the Carboniferous, including early amphibian line ...
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Upper Carboniferous
Upper may refer to: * Shoe upper or ''vamp'', the part of a shoe on the top of the foot * Stimulant Stimulants (also often referred to as psychostimulants or colloquially as uppers) is an overarching term that covers many drugs including those that increase activity of the central nervous system and the body, drugs that are pleasurable and inv ..., drugs which induce temporary improvements in either mental or physical function or both * ''Upper'', the original film title for the 2013 found footage film ''The Upper Footage'' See also

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Late Pennsylvanian
The Pennsylvanian ( , also known as Upper Carboniferous or Late Carboniferous) is, in the ICS geologic timescale, the younger of two subperiods (or upper of two subsystems) of the Carboniferous Period. It lasted from roughly . As with most other geochronologic units, the rock beds that define the Pennsylvanian are well identified, but the exact date of the start and end are uncertain by a few hundred thousand years. The Pennsylvanian is named after the U.S. state of Pennsylvania, where the coal-productive beds of this age are widespread. The division between Pennsylvanian and Mississippian comes from North American stratigraphy. In North America, where the early Carboniferous beds are primarily marine limestones, the Pennsylvanian was in the past treated as a full-fledged geologic period between the Mississippian and the Permian. In parts of Europe, the Mississippian and Pennsylvanian are one more-or-less continuous sequence of lowland continental deposits and are grouped toge ...
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Edaphosaurus Pogonias
''Edaphosaurus'' (, meaning "pavement lizard" for dense clusters of teeth) is a genus of extinct edaphosaurid synapsids that lived in what is now North America and Europe around 303.4 to 272.5 million years ago, during the Late Carboniferous to Early Permian. American paleontologist Edward Drinker Cope first described ''Edaphosaurus'' in 1882, naming it for the "dental pavement" on both the upper and lower jaws, from the Greek ' ("ground"; also "pavement") and (') ("lizard"). ''Edaphosaurus'' is important as one of the earliest-known, large, plant-eating (herbivorous), amniote tetrapods (four-legged land-living vertebrates). In addition to the large tooth plates in its jaws, the most characteristic feature of ''Edaphosaurus'' is a sail on its back. A number of other synapsids from the same time period also have tall dorsal sails, most famously the large apex predator ''Dimetrodon''. However, the sail on ''Edaphosaurus'' is different in shape and morphology. The first fossils o ...
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Sphenacodon Ferox
''Sphenacodon'' (meaning "wedge point tooth") is an extinct genus of synapsid that lived from about 300 to about 280 million years ago (Ma) during the Late Carboniferous and Early Permian periods. Like the closely related ''Dimetrodon'', ''Sphenacodon'' was a carnivorous member of the Eupelycosauria family Sphenacodontidae. However, ''Sphenacodon'' had a low crest along its back, formed from blade-like bones on its vertebrae (neural spines) instead of the tall dorsal sail found in ''Dimetrodon''. Fossils of ''Sphenacodon'' are known from New Mexico and the Utah–Arizona border region in North America. Researchers currently recognize two species: ''Sphenacodon ferox'' (the type species) and ''Sphenacodon ferocior''. ''Sphenacodon ferocior'' can be up to 40% larger in overall size (at about long) compared to ''Sphenacodon ferox'' (at about ). In addition, the dorsal spines in ''Sphenacodon ferocior'' are proportionately 45% taller than in ''Sphenacodon ferox''. The recent discover ...
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