Proarticulata
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Proarticulata
Proarticulata is a proposed phylum of extinct, bilaterally symmetrical animals known from fossils found in the Ediacaran (Vendian) marine deposits, and dates to approximately . The name comes from the Greek () = "before" and Articulata, i.e. prior to animals with true segmentation such as annelids and arthropods. This phylum was established by Mikhail A. Fedonkin in 1985 for such animals as '' Dickinsonia'', ''Vendia'', ''Cephalonega'', ''Praecambridium'' and currently many other Proarticulata are described (see list). Due to their simplistic morphology, their affinities and mode of life are subject to debate. They are almost universally considered to be metazoans, and due to possessing a clear central axis have been suggested to be stem-bilaterians. In the traditional interpretation, the Proarticulatan body is divided into transverse articulation (division) into isomers as distinct from the transverse articulation segments in annelids and arthropods, as their individual isome ...
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Proarticulata
Proarticulata is a proposed phylum of extinct, bilaterally symmetrical animals known from fossils found in the Ediacaran (Vendian) marine deposits, and dates to approximately . The name comes from the Greek () = "before" and Articulata, i.e. prior to animals with true segmentation such as annelids and arthropods. This phylum was established by Mikhail A. Fedonkin in 1985 for such animals as '' Dickinsonia'', ''Vendia'', ''Cephalonega'', ''Praecambridium'' and currently many other Proarticulata are described (see list). Due to their simplistic morphology, their affinities and mode of life are subject to debate. They are almost universally considered to be metazoans, and due to possessing a clear central axis have been suggested to be stem-bilaterians. In the traditional interpretation, the Proarticulatan body is divided into transverse articulation (division) into isomers as distinct from the transverse articulation segments in annelids and arthropods, as their individual isome ...
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Proarticulata Classes
Proarticulata is a proposed phylum of extinct, bilaterally symmetrical animals known from fossils found in the Ediacaran (Vendian) marine deposits, and dates to approximately . The name comes from the Greek () = "before" and Articulata, i.e. prior to animals with true segmentation such as annelids and arthropods. This phylum was established by Mikhail A. Fedonkin in 1985 for such animals as '' Dickinsonia'', '' Vendia'', ''Cephalonega'', ''Praecambridium'' and currently many other Proarticulata are described (see list). Due to their simplistic morphology, their affinities and mode of life are subject to debate. They are almost universally considered to be metazoans, and due to possessing a clear central axis have been suggested to be stem-bilaterians. In the traditional interpretation, the Proarticulatan body is divided into transverse articulation (division) into isomers as distinct from the transverse articulation segments in annelids and arthropods, as their individual i ...
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Isomer (Proarticulata)
Isomer (Greek ''isos'' = "equal", ''méros'' = "part") is an element of transverse body articulation of the bilateral fossil animals of the Phylum Proarticulata from the Ediacaran (Vendian) period. This term has been proposed by Andrey Yu. Ivantsov, a Russian paleontologist from the Laboratory of the Precambrian organisms, Paleontological Institute, Russian Academy of Sciences. Morphology Proarticulatan isomers are distinct from the segments of the Annelida and Panarthropoda, as each of these elements occupies only half of width of a body and are organized in an alternating pattern relatively to the axis of the body. In other words, although proarticulatans are bilaterally symmetrical, one side is not the direct mirror image of its opposite. Opposite isomers of left and right side are located with displacement of half of its width. This phenomenon is described as the symmetry of gliding reflection.M. A. Fedonkin (1985). "Systematic Description of Vendian Metazoa". In Sokolov, ...
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Yorgia
''Yorgia waggoneri'' is a discoid Ediacaran organism. It has a low, segmented body consisting of a short wide "head", no appendages, and a long body region, reaching a maximum length of . It is classified within the extinct animal phylum Proarticulata. Etymology The generic name ''Yorgia'' comes from the Yorga river on the Zimnii Bereg (Winter Coast) of the White Sea, where the first specimens were found. The specific name ''Yorgia waggoneri'' honors the American paleontologist Ben Waggoner, who found the first specimen. Morphology The body plan of the ''Yorgia'' and other proarticulates is unusual for solitary (non-colonial) metazoans. These bilateral organisms have segmented metameric bodies, but left and right transverse elements (isomers) are organized in an alternating pattern relatively to the axis of the body – they are not direct mirror images. This phenomenon is described as the symmetry of glide reflection, which is a characteristic also found in the similar ''Spri ...
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Dickinsonia
''Dickinsonia'' is an extinct genus of basal animal that lived during the late Ediacaran period in what is now Australia, China, Russia and Ukraine. The individual ''Dickinsonia'' typically resembles a bilaterally symmetrical ribbed oval. Its affinities are presently unknown; its mode of growth is consistent with a stem-group bilaterian affinity, though some have suggested that it belongs to the fungi, or even an "extinct kingdom". It lived during the late Ediacaran (part of Precambrian). The discovery of cholesterol molecules in fossils of ''Dickinsonia'' lends support to the idea that ''Dickinsonia'' was an animal. Description ''Dickinsonia'' fossils are known only in the form of imprints and casts in sandstone beds. The specimens found range from a few millimetres to about in length, and from a fraction of a millimetre to a few millimetres thick. They are nearly bilaterally symmetric, segmented, round or oval in outline, slightly expanded to one end (i.e. egg-shaped out ...
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Cephalonega
''Cephalonega stepanovi'' is a fossil organism from Ediacaran deposits of the Arkhangelsk Region, Russia. It was described by Mikhail A. Fedonkin in 1976 Name Its original genus name ''Onega'' comes from the Onega Peninsula of the White Sea, where the first fossils were found. The species name was given to honour V.A. Stepanov, who discovered the Ediacaran fossil site on the Letniy Bereg (" Summer Coast") in 1972, on the Onega Peninsula, the first Proterozoic site found in the Arkhangelsk Oblast. The original generic name is previously occupied by the hemipteran genus ''Onega'' Distant (1908). Ivantsov ''et al.'' (2019) coined a replacement generic name ''Cephalonega''. Morphology The small fossils, which range up to long, have oval outlines and low bodies with an articulated central zone built of isomers encircled by an undivided zone. The surface of the undivided region of ''Cephalonega'' is covered with small tubercles. ''Cephalonega'' was originally described by ...
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Vendiamorpha
Vendiamorpha is a class of extinct animals within the Ediacaran phylum Proarticulata. The typical vendiamorph had an oval-shaped or round-shaped body divided completely into segmented isomers, that were arranged alternately in two rows with reference to the longitudinal axis of the body. Description The phenomenon of left-right alternating segments is called ''glide reflection symmetry'', and is a diagnostic feature of proarticulatans. Transverse elements decrease in size from one end to the other and are inclined in the same direction. The larger isomers cover the smaller ones externally and the first isomer is much larger than the rest. Typically, the first few, or largest isomers are fused together to form a headshield-like structure, leading some researchers to have originally considered them to be ancestral or related to arthropods, though, overwhelming evidence of them being proarticulatans have since led researchers to discard this hypothetical relationship. Som ...
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Vendiidae
Vendiamorpha is a class of extinct animals within the Ediacaran phylum Proarticulata. The typical vendiamorph had an oval-shaped or round-shaped body divided completely into segmented isomers, that were arranged alternately in two rows with reference to the longitudinal axis of the body. Description The phenomenon of left-right alternating segments is called ''glide reflection symmetry'', and is a diagnostic feature of proarticulatans. Transverse elements decrease in size from one end to the other and are inclined in the same direction. The larger isomers cover the smaller ones externally and the first isomer is much larger than the rest. Typically, the first few, or largest isomers are fused together to form a headshield-like structure, leading some researchers to have originally considered them to be ancestral or related to arthropods, though, overwhelming evidence of them being proarticulatans have since led researchers to discard this hypothetical relationship. Som ...
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Cephalozoa
Cephalozoa are an extinct class of primitive segmented marine organisms within the Phylum Proarticulata from the Ediacaran period. They possessed bilateral symmetry and were characterized by a thin, rounded body. Description Unlike the other classes of proarticulates, the segmentation of the body is not complete and shows a "head" with fine distribution channels. Some species of the Yorgiidae family also show some asymmetry.Ivantsov, A. Y. (2004"Vendian Animals in the Phylum Proarticulata" The Rise and Fall of the Vendian Biota. IGSP Project 493. Abstracts. Prato, Italy, . They were discovered in Russia near the White Sea in the Arkhangelsk region, where they lived during the Ediacaran, approximately 635 to 540 Ma (millions of years ago). Taxonomy Cephalozoa includes the families Yorgiidae and Sprigginidae: Yorgiidae *† ''Archaeaspinus'' Ivantsov, 2007 (synonym of ''Archaeaspis'') **† ''Archaeaspinus fedonkini'' Ivantsov, 2001 *† ''Yorgia'' Ivantsov, 1999 **† ''Yorgi ...
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Karakhtia
''Karakhtia nessovi'' is a species of Proarticulate from the Ediacaran period, around 555 Million Years Ago. K. nessovi is the only species in the genus ''Karakhtia''. The genus Haootia has been compared minorly to ''Karakhtia'' in the way that the fossils of ''Haootia'' superficially resemble the crumpled margins of ''Karakhtia''. See also * Vendiamorpha * Vendia * List of ediacaran genera This is a list of all described Ediacaran genera, including the Ediacaran biota. It contains 227 genera. References {{reflist, 30em * Ediacaran The Ediacaran Period ( ) is a geological period that spans 96 million years from the end ... References Vendiamorpha Ediacaran life Ediacaran Proarticulata Enigmatic prehistoric animal genera Taxa described in 2004 {{Ediacaran-stub ...
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Bilaterians
The Bilateria or bilaterians are animals with bilateral symmetry as an embryo, i.e. having a left and a right side that are mirror images of each other. This also means they have a head and a tail (anterior-posterior axis) as well as a belly and a back (ventral-dorsal axis). Nearly all are bilaterally symmetrical as adults as well; the most notable exception is the echinoderms, which achieve secondary pentaradial symmetry as adults, but are bilaterally symmetrical during embryonic development. Most animals are bilaterians, excluding sponges, ctenophores, placozoans and cnidarians. For the most part, bilateral embryos are triploblastic, having three germ layers: endoderm, mesoderm, and ectoderm. Except for a few phyla (i.e. flatworms and gnathostomulids), bilaterians have complete digestive tracts with a separate mouth and anus. Some bilaterians lack body cavities ( acoelomates, i.e. Platyhelminthes, Gastrotricha and Gnathostomulida), while others display primary body cavi ...
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Vendia
''Vendia'' is a genus of oval-shaped, Ediacaran fossils ranging from 4.5 to 12.5 mm long. The body is completely segmented into isomers, which are arranged alternately in two rows longitudinal to the axis of the body. The larger isomers cover the smaller ones externally but the posterior ends of all the isomers remain free. The transverse elements decrease in size from anterior to posterior and are all inclined in the same direction. The fossil bears a depression along a body that is interpreted as a digestive-distributive system that consists of a simple axial tube and short lateral appendages located along the borders between the isomers. Except for the first isomer of ''Vendia rachiata'', all the isomers have one lateral appendage. The first species, ''V. sokolovi'', was originally found in a core from a Yarensk borehole in the south of Arkhangelsk Oblast of Russia in beginning of the 1960s V. V. Menner. (1963). "The Other Problematical Organic Remains". In: "Stratigrap ...
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