Polyodontidae
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Polyodontidae
Paddlefish (family Polyodontidae) are a family of ray-finned fish belonging to order Acipenseriformes, and one of two living groups of the order alongside sturgeons (Acipenseridae). They are distinguished from other fish by their titular elonglated rostrums, which are thought to enhance electroreception to detect prey. Paddlefish have been referred to as "primitive fish" because Acipenseriformes are amongst the earliest diverging lineages of ray-finned fish, having diverged from all other living groups over 300 million years ago. Paddlefish are almost exclusively North American and Chinese, both extant and in the fossil record. There are six known species: four extinct species known only from fossil remains (three from western North America, one from China), one extant species, the American paddlefish (''Polyodon spathula'') which is native to the Mississippi River basin in the U.S., and the Chinese paddlefish (''Psephurus gladius''), declared extinct in 2022 following a 2019 r ...
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Chinese Paddlefish
The Chinese paddlefish (''Psephurus gladius''; : literal translation: "white sturgeon"), also known as the Chinese swordfish, is an Extinction, extinct species of fish that was formerly native to the Yangtze and Yellow River basins in China. With records of specimens over and possibly in length, it was one of the largest species of Fresh water, freshwater fish. It was the only species in the genus ''Psephurus'' and one of two Holocene, recent species of paddlefish (Polyodontidae), the other being the American paddlefish (''Polyodon spathula''). It was an anadromous species, meaning that it spent part of its adult life at sea, while migrating upriver to Spawn (biology), spawn. The Chinese paddlefish was officially declared extinct in 2022, with an estimated time of extinction to be by 2005, and no later than 2010, although it had become functionally extinct by 1993. The main cause of its extinction was the construction of the Gezhouba Dam, Gezhouba and Three Gorges Dam, Three Go ...
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American Paddlefish
The American paddlefish (''Polyodon spathula,'' also known as a Mississippi paddlefish, spoon-billed cat, or spoonbill) is a species of ray-finned fish. It is the only living species of paddlefish (Polyodontidae). This family is most closely related to the sturgeons; together they make up the order Acipenseriformes, which are one of the most primitive living groups of ray-finned fish. Fossil records of other paddlefish species date back 125 million years to the Early Cretaceous, with records of ''Polyodon'' extending back 65 million years to the early Paleocene. The American paddlefish is a smooth-skinned freshwater fish with an almost entirely cartilaginous skeleton and a paddle-shaped rostrum (snout), which extends nearly one-third its body length. It has been referred to as a freshwater shark because of its heterocercal tail or caudal fin resembling that of sharks, though it is not closely related. The American paddlefish is a highly derived fish because it has evolved spec ...
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Acipenseriformes
Acipenseriformes is an order of basal ray-finned fishes that includes living and fossil sturgeons and paddlefishes (Acipenseroidei), as well as the extinct families Chondrosteidae and Peipiaosteidae. They are the second earliest diverging group of living ray-finned fish after the bichirs. Despite being early diverging, they are highly derived, having only weakly ossified skeletons that are mostly made of cartilage, and in modern representatives highly modified skulls. Description The axial skeleton of Acipenseriformes is only partially ossified, with the majority of the bones being replaced with cartilage. The notochord, usually only found in fish embryos, is unconstricted and retained throughout life. The premaxilla and maxilla bones of the skull present in other vertebrates have been lost. The infraorbital nerve is carried by a series of separate canals, rather than being within the circumorbital bones. The palatoquadrate bones of the skull possess a cartilagi ...
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Crossopholis
''Crossopholis'' is an extinct bony fish known from the early Eocene (Ypresian) of North America, approximately 52 million years ago. It is a close relative of the contemporary American paddlefish, belonging to the paddlefish family. History of discovery First described by Edward Drinker Cope in 1883, the first specimen consisted of an incomplete section of the fish's body and tail. In 1886, a partial skull was recovered by Cope. A nearly complete fossil wasn't recorded until 1980. This was due, in part, to the comparative rarity of the fossil as well as the similarities to other species found within the site. Classification A member of the family Polyodontidae, ''Crossopholis'' is most closely related to the American paddlefish. ''Crossopholis'' means "fringed scales"; a reference to the thousands of tiny (less than 0.5mm) scales which covered the body of the animal. Relationships of recent and fossil paddlefish genera, after Grande ''et al.'' (2002). Paleobiology The fossils ...
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Paleopsephurus
''Palaeopsephurus'' is an extinct genus of paddlefish in the Acipenseriformes family Polyodontidae. At present the genus contains the single species ''Palaeopsephurus wilsoni''. The genus is known primarily from the Late Cretaceous (Maastrichtian) aged Hell Creek Formation of Montana. ''Palaeopsephurus'' is one of two extinct paddlefish genera to have been found in the North American fossil record, with the other genus and species '' Crossopholis magnicaudatus'' having been described and found in the Early Eocene Green River Formation. A third extinct genus ''Protopsephurus'' with the single species '' Protopsephurus liui'' was described from China in 1994. Only two modern paddle fish species are known, ''Polyodon spathula'' in the Mississippi River System of North America, and the extinct ''Psephurus gladius'' in the Yangtze River of China. History and classification The genus was described from three partial specimens currently residing in the collections of the University ...
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Sturgeon
Sturgeon is the common name for the 27 species of fish belonging to the family Acipenseridae. The earliest sturgeon fossils date to the Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ..., and are descended from other, earlier Acipenseriformes, acipenseriform fish, which date back to the Early Jurassic period, some 174 to 201 million years ago. They are one of two living families of the Acipenseriformes alongside paddlefish (Polyodontidae). The family is grouped into four genera: ''Acipenser'' (which is paraphyletic, containing many distantly related sturgeon species), ''Huso'', ''Scaphirhynchus,'' and ''Pseudoscaphirhynchus''. Two species (''Adriatic sturgeon, A. naccarii'' and ''Dabry's sturgeon, A. dabryanus'') may be extinct in the wild, and one (''Syr Darya s ...
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Protopsephurus
''Protopsephurus'' is an extinct genus of paddlefish containing the single species ''Protopsephurus liui,'' known from the Yixian Formation in Liaoning, northern China from the Barremian to Aptian ages of the Early Cretaceous period around 125-120 million years ago. It is up to be one meter in length. It is currently the oldest and most basal paddlefish known. Description The species is known from numerous specimens ranging up to about long. The snout is shorter than that in any other known paddlefish, and is more sturgeon-like. The morphology of the skull roof is also more archaic than any other paddlefish. The axial skeleton is poorly ossified. Like other extinct polyodontids, it also has tiny non-interlocking scales approximately 1 mm in diameter called denticles that cover the trunk, which bear a fringe of spikes. Diet ''Protopsephurus'' is thought to have been piscovorous, feeding on smaller fish. One adult specimen of ''Protopsephurus'' was observed with a specimen of ...
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Benthic
The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "the depths." Organisms living in this zone are called benthos and include microorganisms (e.g., bacteria and fungi) as well as larger invertebrates, such as crustaceans and polychaetes. Organisms here generally live in close relationship with the substrate and many are permanently attached to the bottom. The benthic boundary layer, which includes the bottom layer of water and the uppermost layer of sediment directly influenced by the overlying water, is an integral part of the benthic zone, as it greatly influences the biological activity that takes place there. Examples of contact soil layers include sand bottoms, rocky outcrops, coral, and bay mud. Description Oceans The benthic region of the ocean begins at the shore line (intertidal ...
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Aquatic Feeding Mechanisms
Aquatic feeding mechanisms face a special difficulty as compared to feeding on land, because the density of water is about the same as that of the prey, so the prey tends to be pushed away when the mouth is closed. This problem was first identified by Robert McNeill Alexander. As a result, underwater predators, especially bony fish, have evolved a number of specialized feeding mechanisms, such as filter feeding, ram feeding, suction feeding, protrusion, and pivot feeding. Most underwater predators combine more than one of these basic principles. For example, a typical generalized predator, such as the cod, combines suction with some amount of protrusion and pivot feeding. Suction feeding Suction feeding is a method of ingesting a prey item in fluids by sucking the prey into the predator's mouth. It is a highly coordinated behavior achieved by the dorsal rotation of the dermatocranium, lateral expansion of the suspensorium, and the depression of the lower jaw and hyoid. Suc ...
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Zooplankton
Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by currents in the ocean, or by currents in seas, lakes or rivers. Zooplankton can be contrasted with phytoplankton, which are the plant component of the plankton community ("phyto" comes from the Greek word for ''plant''). Zooplankton are heterotrophic (other-feeding), whereas phytoplankton are autotrophic (self-feeding). This means zooplankton cannot manufacture their own food but must eat other plants or animals instead — in particular they eat phytoplankton. Zooplankton are generally larger than phytoplankton, most are microscopic, but some (such as jellyfish) are macroscopic and can be seen with the naked eye. Many protozoans (single-celled protists that prey on other microscopic life) are zooplankton, including zooflagellates, fo ...
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Electron Microscopy
An electron microscope is a microscope that uses a beam of accelerated electrons as a source of illumination. As the wavelength of an electron can be up to 100,000 times shorter than that of visible light photons, electron microscopes have a higher resolving power than light microscopes and can reveal the structure of smaller objects. A scanning transmission electron microscope has achieved better than 50  pm resolution in annular dark-field imaging mode and magnifications of up to about 10,000,000× whereas most light microscopes are limited by diffraction to about 200  nm resolution and useful magnifications below 2000×. Electron microscopes use shaped magnetic fields to form electron optical lens systems that are analogous to the glass lenses of an optical light microscope. Electron microscopes are used to investigate the ultrastructure of a wide range of biological and inorganic specimens including microorganisms, cells, large molecules, biopsy samples, ...
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