Platyhelminthes
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Platyhelminthes
The flatworms, flat worms, Platyhelminthes, or platyhelminths (from the Greek language, Greek πλατύ, ''platy'', meaning "flat" and ἕλμινς (root: ἑλμινθ-), ''helminth-'', meaning "worm") are a Phylum (biology), phylum of relatively simple bilaterian, Segmentation (biology), unsegmented, soft-bodied invertebrates. Unlike other bilaterians, they are acoelomates (having no coelom, body cavity), and have no specialized circulatory system, circulatory and respiratory system, respiratory organ (anatomy), organs, which restricts them to having flattened shapes that allow oxygen and nutrients to pass through their bodies by diffusion. The digestive cavity has only one opening for both ingestion (intake of nutrients) and egestion (removal of undigested wastes); as a result, the food cannot be processed continuously. In traditional medicinal texts, Platyhelminthes are divided into Turbellaria, which are mostly non-parasitic animals such as planarians, and three entirely p ...
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Turbellaria
The Turbellaria are one of the traditional sub-divisions of the phylum Platyhelminthes (flatworms), and include all the sub-groups that are not exclusively parasitic. There are about 4,500 species, which range from to large freshwater forms more than long or terrestrial species like ''Bipalium kewense'' which can reach in length. All the larger forms are flat with ribbon-like or leaf-like shapes, since their lack of respiratory system, respiratory and circulatory system, circulatory systems means that they have to rely on diffusion for internal transport of metabolites. However, many of the smaller forms are round in cross section. Most are predators, and all live in water or in moist terrestrial environments. Most forms reproduce sexual reproduction, sexually and with few exceptions all are hermaphrodites, simultaneous hermaphrodites. The Acoelomorpha and the genus ''Xenoturbella'' were formerly included in the Turbellaria, but are no longer regarded as Platyhelminthes. All th ...
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Acoelomorpha
Acoelomorpha is a subphylum of very simple and small soft-bodied animals with planula-like features which live in marine or brackish waters. They usually live between grains of sediment, swimming as plankton, or crawling on other organisms, such as algae and corals.Cannon, L. R. G. (1986) ''Turbellaria of the World. A guide to families and genera''. Brisbane, Queensland Museum, 136 p. With the exception of two acoel freshwater species, all known Acoelomorphs are marine. Systematics Etymology The term "acoelomorph" derives from the Ancient Greek words (), the ''alpha privative'', expressing negation or absence, (), meaning "cavity", and (), meaning "form". This refers to the fact that acoelomorphs have a structure lacking a fluid-filled body cavity. Classification The subphylum Acoelomorpha is divided into two classes. There are at least 408 described species, with a majority of these falling within the Crucimusculata infraorder in Acoela. * Acoela comprise small fla ...
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Proglottid
Cestoda is a class of parasitic worms in the flatworm phylum (Platyhelminthes). Most of the species—and the best-known—are those in the subclass Eucestoda; they are ribbon-like worms as adults, known as tapeworms. Their bodies consist of many similar units known as proglottids—essentially packages of eggs which are regularly shed into the environment to infect other organisms. Species of the other subclass, Cestodaria, are mainly fish infecting parasites. All cestodes are parasitic; many have complex life histories, including a stage in a definitive (main) host in which the adults grow and reproduce, often for years, and one or two intermediate stages in which the larvae develop in other hosts. Typically the adults live in the digestive tracts of vertebrates, while the larvae often live in the bodies of other animals, either vertebrates or invertebrates. For example, '' Diphyllobothrium'' has at least two intermediate hosts, a crustacean and then one or more freshwater fi ...
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Cestoda
Cestoda is a class of parasitic worms in the flatworm phylum (Platyhelminthes). Most of the species—and the best-known—are those in the subclass Eucestoda; they are ribbon-like worms as adults, known as tapeworms. Their bodies consist of many similar units known as proglottids—essentially packages of eggs which are regularly shed into the environment to infect other organisms. Species of the other subclass, Cestodaria, are mainly fish infecting parasites. All cestodes are parasitic; many have complex life histories, including a stage in a definitive (main) host in which the adults grow and reproduce, often for years, and one or two intermediate stages in which the larvae develop in other hosts. Typically the adults live in the digestive tracts of vertebrates, while the larvae often live in the bodies of other animals, either vertebrates or invertebrates. For example, '' Diphyllobothrium'' has at least two intermediate hosts, a crustacean and then one or more freshwater fi ...
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Catenulida
Catenulida is an order of flatworms in the classical classification, or a class of flatworms in a phylogenetic approach. They are relatively small free-living flatworms, inhabiting freshwater and marine environments. There are about 100 species described worldwide, but the simple anatomy makes species distinction problematic. Description The anatomy of catenulids is simple and lacks hard parts. The mouth is located anteriorly and connects to a simple pharynx and a simple intestine that forms a ciliated sac. They possess two pairs of nerve cords and often a statocyst, as well as a single protonephridium. The gonads are unpaired. Unusually, the male gonopore opens on the dorsal surface of the animal, above the pharynx, while the female reproductive system lacks any of the usual ducts and related structures found in other flatworms. The sperm is nonmobile and lacks flagella or cilia. Asexual reproduction by paratomy is common, and it usually leads to a chain of organisms (zooids), ...
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Invertebrate
Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate subphylum Vertebrata. Familiar examples of invertebrates include arthropods, mollusks, annelids, echinoderms and cnidarians. The majority of animal species are invertebrates; one estimate puts the figure at 97%. Many invertebrate taxa have a greater number and variety of species than the entire subphylum of Vertebrata. Invertebrates vary widely in size, from 50  μm (0.002 in) rotifers to the 9–10 m (30–33 ft) colossal squid. Some so-called invertebrates, such as the Tunicata and Cephalochordata, are more closely related to vertebrates than to other invertebrates. This makes the invertebrates paraphyletic, so the term has little meaning in taxonomy. Etymology The word "invertebrate" comes from the Latin word ''vertebra'', whi ...
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Bilaterian
The Bilateria or bilaterians are animals with bilateral symmetry as an embryo, i.e. having a left and a right side that are mirror images of each other. This also means they have a head and a tail (anterior-posterior axis) as well as a belly and a back (ventral-dorsal axis). Nearly all are bilaterally symmetrical as adults as well; the most notable exception is the echinoderms, which achieve secondary pentaradial symmetry as adults, but are bilaterally symmetrical during embryonic development. Most animals are bilaterians, excluding sponges, ctenophores, placozoans and cnidarians. For the most part, bilateral embryos are triploblastic, having three germ layers: endoderm, mesoderm, and ectoderm. Except for a few phyla (i.e. flatworms and gnathostomulids), bilaterians have complete digestive tracts with a separate mouth and anus. Some bilaterians lack body cavities (Acoelomata, acoelomates, i.e. Flatworm, Platyhelminthes, Gastrotricha and Gnathostomulida), while others display pri ...
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Rhabditophora
Rhabditophora (from ''rhabdito''-, rhabdite + Greek -φορος ''phoros'' bearer, i.e., "rhabdite bearers") is a class of flatworms. It includes all parasitic flatworms (clade Neodermata) and most free-living species that were previously grouped in the now obsolete class Turbellaria. Therefore, it contains the majority of the species in the phylum Platyhelminthes, excluding only the catenulids, to which they appear to be the sister group. The clade Rhabditophora was originally erected by Ulrich Ehlers in 1985Ehlers, U. (1985) ''Phylogenetic relationships within the Platyhelminthes''. ''In'' S. Conway Morris; J. D. George; R. Gibson; H. M. Platt (Eds.), ''The origins and relationships of lower invertebrates''. Oxford, Clarendon Press, p. 143-158. based on morphological analyses and its monophyly was later confirmed by molecular studies. Description Rhabditophorans are characterized by the presence of lamellated rhabdites, rodlike granules secreted in the cells of the epidermis ...
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Planarian
A planarian is one of the many flatworms of the traditional class Turbellaria. It usually describes free-living flatworms of the order Tricladida (triclads), although this common name is also used for a wide number of free-living platyhelminthes. Planaria are common to many parts of the world, living in both saltwater and freshwater ponds and rivers. Some species are terrestrial and are found under logs, in or on the soil, and on plants in humid areas. The triclads are characterized by triply branched intestine and anteriorly situated ovaries, next to the brain. Today the order Tricladida is split into three suborders, according to their phylogenetic relationships: Maricola, Cavernicola and Continenticola. Formerly, the Tricladida was split according to habitats: Maricola, which is marine; Paludicola which inhabits freshwater; and Terricola, which is land-dwelling. Planaria exhibit an extraordinary ability to regenerate lost body parts. For example, a planarian split lengt ...
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Lophotrochozoa
Lophotrochozoa (, "crest/wheel animals") is a clade of protostome animals within the Spiralia. The taxon was established as a monophyletic group based on molecular evidence. The clade includes animals like annelids, molluscs, bryozoans, brachiopods, and platyhelminthes. Groups Lophotrochozoa was defined in 1995 as the "last common ancestor of the three traditional lophophorate taxa (brachiopods, bryozoans, and phoronid worms), the mollusks and the annelids, and all of the descendants of that common ancestor". It is a cladistic definition (a node-based name), so the affiliation to Lophotrochozoa of spiralian groups not mentioned directly in the definition depends on the topology of the spiralian tree of life, and in some phylogenetic hypotheses, Lophotrochozoa may even be synonymous to Spiralia. Nemertea and Orthonectida (if not directly considered as part of Annelida) are probably lophotrochozoan phyla; Dicyemida, Gastrotricha, and Platyhelminthes may be lophotrochozoans ...
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Monogenea
Monogeneans are a group of ectoparasitic flatworms commonly found on the skin, gills, or fins of fish. They have a direct lifecycle and do not require an intermediate host. Adults are hermaphrodites, meaning they have both male and female reproductive structures.L.A. Tubbsa et al. (2005). "Effects of temperature on fecundity in vitro, egg hatching and reproductive development of ''Benedenia seriolae'' and ''Zeuxapta seriolae'' (Monogenea) parasitic on yellowtail kingfish Seriola lalandi". ''International Journal for Parasitology''(35), 315–327. Some monogeneans are oviparous (egg-laying) and some are viviparous (live-bearing). Oviparous varieties release eggs into the water. Viviparous varieties release larvae, which immediately attach to another host. The genus ''Gyrodactylus'' is an example of a viviparous variety, while the genus ''Dactylogyrus'' is an example of an oviparous variety. Signs and symptoms Freshwater fish that become infected with this parasite become let ...
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Phylum (biology)
In biology, a phylum (; plural: phyla) is a level of classification or taxonomic rank below kingdom and above class. Traditionally, in botany the term division has been used instead of phylum, although the International Code of Nomenclature for algae, fungi, and plants accepts the terms as equivalent. Depending on definitions, the animal kingdom Animalia contains about 31 phyla, the plant kingdom Plantae contains about 14 phyla, and the fungus kingdom Fungi contains about 8 phyla. Current research in phylogenetics is uncovering the relationships between phyla, which are contained in larger clades, like Ecdysozoa and Embryophyta. General description The term phylum was coined in 1866 by Ernst Haeckel from the Greek (, "race, stock"), related to (, "tribe, clan"). Haeckel noted that species constantly evolved into new species that seemed to retain few consistent features among themselves and therefore few features that distinguished them as a group ("a self-contained uni ...
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