Neoceratopsians
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Neoceratopsians
Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassic. The earliest known ceratopsian, ''Yinlong downsi'', lived between 161.2 and 155.7 million years ago.Holtz, Thomas R. Jr. (2011) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,'Winter 2010 Appendix./ref> The last ceratopsian species, ''Triceratops prorsus'', became extinct during the Cretaceous–Paleogene extinction event, . ''Triceratops'' is by far the best-known ceratopsian to the general public. It is traditional for ceratopsian genus names to end in "''-ceratops''", although this is not always the case. One of the first named genera was ''Ceratops'' itself, which lent its name to the group, although it is considered a ''nomen dubium'' today as its fossil remains have no distinguishing characte ...
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Mosaiceratops
''Mosaiceratops'' is a genus of ceratopsian, described by Zheng, Jin & Xu in 2015 and found in the Xiaguan Formation of Neixiang County. ''Mosaiceratops'' lived in the upper Cretaceous in what is now the Henan Province of China. Although phylogenetic analyses have found ''Mosaiceratops'' to be the most basal neoceratopsian, the authors noted that several features in the premaxilla and nasal bones are shared with ''Psittacosaurus'', indicating that neoceratopsians evolved premaxillary teeth twice and that ''Psittacosaurus'' is not as primitive as previously thought.Zheng, W., Jin, X., & Xu, X. (2015)A psittacosaurid-like basal neoceratopsian from the Upper Cretaceous of central China and its implications for basal ceratopsian evolution ''Scientific reports'', 5. article number 14190: 1-9; doi:10.1038/srep14190 Discovery On the westbank of the river Tuanhe in Neixiang in Henan the skeleton was discovered of a small ceratopian. The fossil was prepared by Sheng Yiming and Yu Chaoh ...
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Liaoceratops
''Liaoceratops'', meaning "Liaoning horned face", is a ceratopsian dinosaur believed to be an early relative of the horned ceratopsids. It lived in the Early Cretaceous, 126 million years ago. It was discovered in China by a team of American and Chinese scientists. ''Liaoceratops'' was much smaller than its later relatives, but offers a glimpse into the early evolution of this group of dinosaurs. Discoveries and naming ''Liaoceratops'' was discovered in the famous Liaoning Province of China, where several fossils of feathered dinosaurs have also been collected. The type species ''Liaoceratops yanzigouensis'' was in 2002 named and described by Xu Xing, Peter Makovicky, Wang Xiaolin, Mark Norell and You Hailu. The generic name is derived from Liaoning and the Greek ''keras'', "horn" and ''ops'', "face". The specific name refers to the town Yanzigou. The holotype IVPP V12738 has been found in the Yixian Formation dating from the Barremian. These beds have also yielded fossil ...
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Beg Tse
''Beg'' (after the Himalayan war deity Beg-tse) is a genus of neoceratopsian dinosaur from the early Cretaceous period of Mongolia. The genus contains a single species, ''Beg tse'', known from a partial skull and very fragmentary postcrania. ''Beg'' represents the most basal neoceratopsian currently known. Discovery and naming The holotype, IGM 100/3652, was discovered in 2015 near the town of Tsogt-Ovoo in the Ömnögovi Province of Mongolia. Described from the Ulaanoosh Formation, the specimen is dated to 113 to 94 million years ago, at the boundary of the Lower and Upper Cretaceous. ''Beg'' is named after Beg-tse, a Himalayan deity who is the god of war in Mongolian culture. The deity is often depicted with a rugose face and/or body, similar to the appearance of the preserved skull of the dinosaur. Description Based on the size of the skull, about long, ''Beg'' was most likely a medium-sized basal ceratopsian, similar in size to ''Yinlong'' and ''Liaoceratops.'' ...
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Auroraceratops
''Auroraceratops'', meaning "dawn horned face", is a genus of bipedal basal neoceratopsian dinosaur, from the Early Cretaceous (Aptian age) of north central China. The etymology of the generic name refers to its status as an early ceratopsian and also to Dawn Dodson, wife of Peter Dodson, one of the palaeontologists who described it. Discovery and species The first specimen was found in the Xinminpu Group, Gongpoquan Basin of the Mazong Shan area of Gansu Province, north central China. As of 2021, fossils from more than 80 more individuals have been found, including complete skeletons. The specific descriptor, "''rugosus''", Latin for "rough", refers to the various rugose areas on the surface of both skull and jaw, namely the wrinkled expansion of the lacrimal bone distinctive of this animal. ''Auroraceratops'' is the second basal neoceratopsian to be found in the Mazong Shan area, after ''Archaeoceratops''. The type specimen, IG-2004-VD-001, consists of a nearly complete ...
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Aquilops
''Aquilops'' is an early herbivorous ceratopsian dinosaur dating from the Early Cretaceous of North America, approximately 109 million to 104 million years ago. The type species is ''A. americanus''. Description The skull is 84.2 mm long. The holotype is possibly not from a full-grown individual. A comparison with related species indicates it might have been at 60% of its adult length. Wedel estimated the total body length of ''Aquilops'' at 60 cm and its weight at 1.5 kg. The authors established some unique traits. The rostral, the bone core of the snout beak, curves downwards and has an arched keel on its top with a bump on the front. In front of the tooth row the upper jaw rim is over its total length concave in side view. The skull opening, the antorbital fenestra, is twice as long as it is tall and has a pointed rear, below the eye socket. Discovery and naming In 1997, paleontologist Scott Madsen found the single fossil, a partial skull, in Carbon County ...
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Helioceratops
''Helioceratops'' is a genus of herbivorous neoceratopsian dinosaur from the Middle Cretaceous of China. Discovery and naming In 2000 and 2002, at the Liufangzi site of China's eastern Jilin province, excavations took place during which the jaws were found of a ceratopsian new to science. The type species ''Helioceratops brachygnathus'' was named and described in 2009 by Jin Liyong, Chen Jun, Zan Shuqin and Pascal Godefroit. The generic name means "sun horned face" from the Greek ''helios'', "sun", ''keras'', "horn" and ''ops'', "face". The reference is that the Sun rises in the East and ceratopsians also "rose" in the East; i.e. they originated in the Orient. The name also refers to a close relationship with '' Auroraceratops'', the "dawn ceratopian". The specific name means "short jaw" from the Greek βραχύς, ''brachys'' and γνάθος, ''gnathos'', in reference to the distinguishing short lower jaw. ''Helioceratops'' was discovered in a layer of the Quantou Forma ...
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Leptoceratopsidae
Leptoceratopsidae is an extinct family of neoceratopsian dinosaurs from Asia, North America and Europe. Leptoceratopsids resembled, and were closely related to, other neoceratopsians, such as the families Protoceratopsidae and Ceratopsidae, but they were more primitive and generally smaller. Phylogeny Leptoceratopsidae was originally named by Franz Nopcsa von Felső-Szilvás in 1923 as a subfamily Leptoceratopsinae, and its type species is ''Leptoceratops gracilis''. Mackovicky, in 2001, defined it as a stem-based taxon and a family consisting of ''Leptoceratops gracilis'' and all species closer to ''Leptoceratops'' than to ''Triceratops horridus''.Makovicky, P.J. 2001. A ''Montanoceratops cerorhynchus'' (Dinosauria: Ceratopsia) braincase from the Horseshoe Canyon Formation of Alberta, In: Tanke, D.H. & Carpenter, K. (Eds.). ''Mesozoic Vertebrate Life''. Bloomington: Indiana University Press. Pp. 243-262. The cladogram below follows the topologies from a 2015 analysis by Yiming ...
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Yamaceratops
''Yamaceratops'' is a genus of primitive ceratopsian that lived in Asia during the Late Cretaceous period in what is now the Javkhlant Formation. Initially, the rocks where it was found in were thought to be from the Early Cretaceous, but the age was reevaluated in 2009. It was a relatively small dinosaur, reaching in length and in body mass. The type species, ''Yamaceratops dorngobiensis'', was described by P. J. Makovicky and M. A. Norell in September, 2006. The authors consider the animal to have had an intermediate phylogenetic position between ''Liaoceratops'' and '' Archaeoceratops'' within Neoceratopia. Examination of the frill of ''Yamaceratops'' has convinced the authors that the frill was not used for display, and that the fossils " intat a more complex evolutionary history for ceratopsian frills." The genus name refers to Yama, a Tibetan Buddhist deity; the species name to the Eastern Gobi. The holotype IGM 100/1315 consists of a partial skull; other material h ...
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Protoceratopsidae
Protoceratopsidae is a family of basal (primitive) ceratopsians from the Late Cretaceous period. Although ceratopsians have been found all over the world, protoceratopsids are only definitively known from Cretaceous strata in Asia, with most specimens found in China and Mongolia. As ceratopsians, protoceratopsids were herbivorous, with constantly replacing tooth batteries made for slicing through plants and a hooked beak for grabbing them. Protoceratopsids were small ceratopsians around 1-2.5 m in length. Their bony frill and horns were much smaller than more derived members of Ceratopsia, such as ceratopsids. Description Protoceratopsids were relatively small ceratopsians, averaging around 1-2.5 m in length from head to tail. Protoceratopsids have a frill and rostral bone characteristic of all ceratopsians. Their snout is particularly wedge-shaped with tall and narrow nostrils situated high on it. The antorbital fenestra is unusually small, and the antorbital fossa sits high on ...
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Archaeoceratops
''Archaeoceratops'', meaning "ancient horned face", is a genus of basal neoceratopsian dinosaur from the Early Cretaceous (Aptian stage) of north central China. It appears to have been bipedal and quite small, reaching in length and in body mass. It had a comparatively large head but no horns, possessing only a small bony frill projecting from the back of its head. Discovery and species Two specimens were found in the Middle Gray Unit of the Xinminbao Group, Gongpoquan Basin of the Mazong Shan area of Gansu Province, north central China. The type species, ''A. oshimai'', was named by Dong Zhiming and Azuma in 1997. It is the first basal neoceratopsian discovered in this area. The type specimen, IVPP V11114, consists of a partially complete skeleton including skull, caudal vertebrae, pelvis, and most of a hind foot. The second specimen (paratype), IVPP V11115, consists of an incomplete skeleton with a relatively well preserved caudal series, a partial hind limb, and a co ...
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Ajkaceratops
''Ajkaceratops'' (pronounced "oi-ka-sera-tops") is a genus of ceratopsian dinosaur described in 2010. It lived during the Late Cretaceous in the western Tethyan archipelago, in what is now Europe. The type species, ''A. kozmai'', is most closely related to forms in east Asia, from where its ancestors may have migrated by island-hopping. The generic name, ''Ajkaceratops'', honors Ajka, a town in Hungary where the fossils were first discovered, combined with the given greek nomination ''ceratops'', meaning "horned face". The specific name, "''kozmai"'', honors Károly Kozma. Description The holotype, cataloged as MTM V2009.192.1, consists only of a few skull fragments, including rostral bones, fused premaxillae, and maxillae fragments (beak and jaw fragments). These fossils are kept in the Hungarian Natural History Museum, in Budapest. Although the fossils are fragmentary, the paper describing ''Ajkaceratops'' estimated a body length of . Other material includes four predentary ...
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Turanoceratops
''Turanoceratops'' ("Turan horned face") is a genus of herbivorous ceratopsian dinosaur from the late Cretaceous Bissekty Formation of Uzbekistan. The fossils dated from the mid-late Turonian stage, roughly 90 million years ago. The skull bore a pair of long brow horns like those seen in the Ceratopsidae, although ''Turanoceratops'' appears to have been transitional between earlier ceratopsians and ceratopsids, and not a ceratopsid itself. Discovery and naming From the 1920s onwards, Soviet scientists discovered fragmentary fossils near Dzharakuduk in the district Navoi Viloyat, leading them to the conclusion that some ceratopsid must have been present. In 1988, paleontologist Lev Aleksandrovich Nesov based on these published the name ''Turanoceratops tardabilis'', but did not provide a description so that for the time being it remained a ''nomen nudum''. In 1989, Nesov, L.F. Kaznysjkina and Gennady Olegovich Cherepanov validly named the type species ''Turanoceratops tardabilis''. ...
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