Holocephali
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Holocephali
Holocephali ("complete heads"), sometimes given the term Euchondrocephali, is a subclass of cartilaginous fish in the class Chondrichthyes. The earliest fossils are of teeth and come from the Devonian period. Little is known about these primitive forms, and the only surviving group in the subclass is the order Chimaeriformes. Chimaeriformes, commonly known as chimaeras, includes the rat fishes in the genus '' Chimaera'', and the elephant fishes in the genus ''Callorhinchus''. These fishes move by using sweeping movements of their large pectoral fins. They are deep sea fish with slender tails, living close to the seabed to feed on benthic invertebrates. They lack a stomach, their food moving directly into the intestine. Extinct holocephalans were much more diverse in lifestyles, including shark-like predatory forms and slow, durophagous fish. Characteristics Members of this taxon preserve today some features of elasmobranch life in Paleozoic times, though in other respects th ...
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Chondrichthyes
Chondrichthyes (; ) is a class that contains the cartilaginous fishes that have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or ''bony fishes'', which have skeletons primarily composed of bone tissue. Chondrichthyes are jawed vertebrates with paired fins, paired nares, scales, and a heart with its chambers in series. Extant chondrichthyes range in size from the 10 cm (3.9 in) finless sleeper ray to the 10 m (32 ft) whale shark. The class is divided into two subclasses: Elasmobranchii (sharks, rays, skates, and sawfish) and Holocephali ( chimaeras, sometimes called ghost sharks, which are sometimes separated into their own class). Within the infraphylum Gnathostomata, cartilaginous fishes are distinct from all other jawed vertebrates. Anatomy Skeleton The skeleton is cartilaginous. The notochord is gradually replaced by a vertebral column during development, except in Holocephali, where the notochord stays intact. In some deepwat ...
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Dwykaselachus
''Dwykaselachus'' (pronounced dwike-a-selak-us) is an extinct genus of symmoriiform, a cartilaginous fish that lived in what is now South Africa during the Permian period around 280 million years ago. It was first discovered in the 1980s, in a nodule of sediments from the Karoo Supergroup. ''Dwykaselachus'' was named based on Dwyka Group, the group of sedimentary geological formation in the southeastern part of Africa. It represents the place where the type species ''Dwykaselachus oosthuizeni'' was found. Prior to its discovery, symmoriiforms were thought to be related to sharks, in the group Elasmobranchii. However, CT scans of its relatively intact skull showed traits such as brain shape and inner ear structure that are shared with cartilaginous fish from the group Holocephali, which includes Chimaera, chimaeras. This implies that the first major radiation of cartilaginous fish after the Late Devonian extinction, Devonian extinction was in fact holocephalians, rather than shar ...
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Chimaera (genus)
''Chimaera'' is the type genus of the cartilaginous fish family Chimaeridae. Species There are currently 16 recognized species in this genus: * '' Chimaera argiloba'' Last, W. T. White & Pogonoski, 2008 (Whitefin chimaera) * '' Chimaera bahamaensis'' Kemper, Ebert, Didier & Compagno, 2010 (Bahamas ghost shark) * ''Chimaera carophila'' Kemper, Ebert, Naylor & Didier, 2014 (Brown chimaera) * ''Chimaera cubana'' Howell-Rivero, 1936 (Cuban chimaera) * ''Chimaera fulva'' Didier, Last & W. T. White, 2008 (Southern chimaera) * ''Chimaera jordani'' S. Tanaka (I), 1905 (Jordan's chimaera) * ''Chimaera lignaria'' Didier, 2002 (Carpenter's chimaera) * ''Chimaera macrospina'' Didier, Last & W. T. White, 2008 (Longspine chimaera) * ''Chimaera monstrosa'' Linnaeus, 1758 (Rabbit fish) * ''Chimaera notafricana'' Kemper, Ebert, Compagno & Didier, 2010 (Cape chimaera) * ''Chimaera obscura'' Didier, Last & W. T. White, 2008 (Shortspine chimaera) * ''Chimaera opalescens'' L ...
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Durophagous
Durophagy is the eating behavior of animals that consume hard-shelled or exoskeleton bearing organisms, such as corals, shelled mollusks, or crabs. It is mostly used to describe fish, but is also used when describing reptiles, including fossil turtles, placodonts and invertebrates, as well as "bone-crushing" mammalian carnivores such as hyenas. Durophagy requires special adaptions, such as blunt, strong teeth and a heavy jaw. Bite force is necessary to overcome the physical constraints of consuming more durable prey and gain a competitive advantage over other organisms by gaining access to more diverse or exclusive food resources earlier in life. Those with greater bite forces require less time to consume certain prey items as a greater bite force can increase the net rate of energy intake when foraging and enhance fitness in durophagous species. In the order Carnivora there are two dietary categories of durophagy; bonecrackers and bamboo eaters. Bonecrackers are exemplified by hy ...
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Chimaera
Chimaeras are cartilaginous fish in the order Chimaeriformes , known informally as ghost sharks, rat fish, spookfish, or rabbit fish; the last three names are not to be confused with rattails, Opisthoproctidae, or Siganidae, respectively. At one time a "diverse and abundant" group (based on the fossil record), their closest living relatives are sharks and rays, though their last common ancestor with them lived nearly 400 million years ago. Today, they are largely confined to deep water. Description and habits Chimaeras live in temperate ocean floors down to deep, with few occurring at depths shallower than . Exceptions include the members of the genus ''Callorhinchus'', the rabbit fish and the spotted ratfish, which locally or periodically can be found at shallower depths. Consequently, these are also among the few species from the chimaera order kept in public aquaria. They live in all the oceans except for the Arctic and Antarctic oceans. They have elongated, soft ...
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Elasmobranchii
Elasmobranchii () is a subclass of Chondrichthyes or cartilaginous fish, including sharks (superorder Selachii), rays, skates, and sawfish (superorder Batoidea). Members of this subclass are characterised by having five to seven pairs of gill clefts opening individually to the exterior, rigid dorsal fins and small placoid scales on the skin. The teeth are in several series; the upper jaw is not fused to the cranium, and the lower jaw is articulated with the upper. The details of this jaw anatomy vary between species, and help distinguish the different elasmobranch clades. The pelvic fins in males are modified to create claspers for the transfer of sperm. There is no swim bladder; instead, these fish maintain buoyancy with large livers rich in oil. The definition of the clade is unclear with respect to fossil chondrichthyans. It has been used by different authors as equivalent to Neoselachii (the clade including modern sharks and rays and their last common ancestor) or for al ...
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Callorhinchus
''Callorhinchus'', the plough-nosed chimaeras or elephantfish, are the only living genus in the family Callorhinchidae (sometimes spelled Callorhynchidae). A few extinct genera only known from fossil remains are recognized. ''Callorhinchus'' spp, are similar in form and habits to other Chimaeriformes, chimaeras, but are distinguished by the presence of an elongated, flexible, fleshy snout, with a vague resemblance to a ploughshare. They are only found in the oceans of the Southern Hemisphere along the ocean bottom on muddy and sandy substrates. They filter feed, with small shellfish making up the bulk of their diet. The plough-nosed chimaera lays eggs on the ocean floor that hatch at around 8 months. They are currently not a target of conservation efforts; however, they may be susceptible to overfishing and trawling. Morphology Plough-nose chimaeras range from about in total length. Their usual color is black or brown, and, often a mixture between the two. While the club-like sn ...
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Symmoriida
Symmoriiformes is an extinct order of holocephalians. Originally named Symmoriida by Zangerl (1981), it has subsequently been known by several other names. Lund (1986) synonymized the group with Cladodontida, while Maisey (2008) corrected the name to Symmoriiformes in order to prevent it from being mistaken for a family. The symmoriiform fossils record appear at the beginning of the Carboniferous. Most of them died out at the start of the Permian, but ''Dwykaselachus'' is known from the Artinskian-Kungurian of South Africa. Teeth described from the Valanginian of France and Austria indicate that members of the family Falcatidae might have survived until the Early Cretaceous; however, these teeth were also argued to be more likely neoselachian teeth. Fossil distribution Fossil evidence of Symmoriida have been found at Bear Gulch, Fergus County, Montana, Bethel Quarry, Pike County, Indiana, Kinshozan quarry, Alaska, Gifu Prefecture, Japan, Bashkortostan, Russian Federation an ...
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Eugeneodontida
The Eugeneodontida is an extinct and poorly known order of cartilaginous fishes. They possessed "tooth-whorls" on the symphysis of either the lower or both jaws and pectoral fins supported by long radials. They probably lacked pelvic fins and anal fins. The palatoquadrate was either fused to the skull or reduced. Now determined to be within the Holocephali, their closest living relatives are ratfish. The eugeneodonts are named after paleontologist Eugene S. Richardson, Jr. The Eugeneodontida disappeared in the Early Triassic. Members of the Eugeneodontida are further classified into different families, the most well-preserved members that have been discovered are commonly placed within the families Helicoprionidae ("spiral saws"), and Edestidae ("those which devour"), the former containing the genera ''Helicoprion'', '' Sarcoprion'', and '' Parahelicoprion'', and the latter containing the genera '' Edestus'', ''Lestrodus'', and '' Metaxyacanthus''. All eugeneodonts are thought ...
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Iniopterygiformes
Iniopterygiformes ("Nape Wing Forms") is an extinct order of chimaera-like cartilaginous fish that lived from the Devonian to Carboniferous periods (345–280 million years ago). Fossils of them have been found in Montana, Indiana, Illinois, and Nebraska. The Iniopterygians are characterized by large pectoral fins, wing-like projections on their backs, mounted high on the body and denticulated bony plates on the head and jaws. Iniopterygian sharks were small, and their average length was about . The elongated pectoral fins had denticles along the leading edge which may have had a role in mating. They are thought to have been able to move their pectorals in a vertical plane, ”flying” through the water much like modern-day flying fish. The majority of iniopterygians are placed within the family Sibyrhinchidae. Members of this family include '' Sibyrhinchus denisoni'', '' Inioptera richardsoni'', and '' Inioxyele''. Ironically, '' Iniopteryx rushlaui'', the type species In zoo ...
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Helicoprion Bessonovi
''Helicoprion'' is an extinct genus of shark-like eugeneodont fish. Almost all fossil specimens are of spirally arranged clusters of the individuals' teeth, called "tooth whorls", which in life were embedded in the lower jaw. As with most extinct cartilaginous fish, the skeleton is mostly unknown. Fossils of ''Helicoprion'' are known from a 20 million year timespan during the Permian period from the Artinskian stage of the Cisuralian (Early Permian) to the Roadian stage of the Guadalupian (Middle Permian). The closest living relatives of ''Helicoprion'' (and other eugeneodonts) are the chimaeras, though their relationship is very distant. The unusual tooth arrangement is thought to have been an adaption for feeding on soft bodied prey, and may have functioned as a deshelling mechanism for hard bodied cephalopods such as nautiloids and ammonoids. In 2013, systematic revision of ''Helicoprion'' via morphometric analysis of the tooth whorls found only ''H. davisii, H. bessonowi'' an ...
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Permian
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the sauropsids ( reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their am ...
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