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Cladonia Cariosa
''Cladonia cariosa'', the split-peg lichen or the split-peg soldiers cup lichen, is a species of fruticose, cup lichen in the family Cladoniaceae. It was first formally named by Erik Acharius in 1799 as ''Lichen cariosus'' and transferred to the genus ''Cladonia'' in 1827 by Kurt Polycarp Joachim Sprengel Kurt Polycarp Joachim Sprengel (3 August 1766 – 15 March 1833) was a German botanist and physician who published an influential multivolume history of medicine, ''Versuch einer pragmatischen Geschichte der Arzneikunde'' (1792–99 in four vol .... It has a broad distribution, occurring in Europe, Asia, North America, and South America. References cariosa Lichen species Taxa named by Erik Acharius Lichens described in 1799 {{Lecanorales-stub ...
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Turnbull National Wildlife Refuge
The Turnbull National Wildlife Refuge is located six miles (10 km) south of Cheney, Washington, on the eastern edge of the Columbia Basin in Spokane County in northeastern Washington. Turnbull NWR encompasses more than of the Channeled Scablands. The ecosystem that predominates the refuge is unique within the National Wildlife Refuge System and has characteristics that distinguish it from natural reserves worldwide. The combination of basalt outcrops, channeled canyons, and ponderosa pine forests infused in a diverse landscape of over 130 marshes, wetlands, and lakes creates an environment of aesthetic beauty as well as high quality wildlife habitat. The refuge is named for Cyrus Turnbull who lived on the site in the 1880s. Geology The refuge is situated within the Channeled Scablands, an area formed 16,000 years ago by Missoula Floods during the last ice age. The powerful forces of volcanism, glaciation, and the largest flood in geological history have combined to forge a ...
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Washington (state)
Washington (), officially the State of Washington, is a state in the Pacific Northwest region of the Western United States. Named for George Washington—the first U.S. president—the state was formed from the western part of the Washington Territory, which was ceded by the British Empire in 1846, by the Oregon Treaty in the settlement of the Oregon boundary dispute. The state is bordered on the west by the Pacific Ocean, Oregon to the south, Idaho to the east, and the Canadian province of British Columbia to the north. It was admitted to the Union as the 42nd state in 1889. Olympia is the state capital; the state's largest city is Seattle. Washington is often referred to as Washington state to distinguish it from the nation's capital, Washington, D.C. Washington is the 18th-largest state, with an area of , and the 13th-most populous state, with more than 7.7 million people. The majority of Washington's residents live in the Seattle metropolitan area, the center of trans ...
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Erik Acharius
Erik Acharius (10 October 1757 – 14 August 1819) was a Swedish botanist who pioneered the taxonomy of lichens and is known as the "father of lichenology." Acharius was famously the last pupil of Carl Linnaeus. Life Acharius was born in 1757 to Johan Eric Acharius and Catharina Margaretha Hagtorn in Gävle.Sernander., K. “Erik Acharius - Svenskt Biografiskt Lexikon.” Fredrik Teodor Borg - Svenskt Biografiskt Lexikon, sok.riksarkivet.se/sbl/Presentation.aspx?id=5503. He received a private education until he was admitted to Gävle Gymnasium in 1770. Later he matriculated at Uppsala University in 1773 where he studied natural history and medicine under Linnaeus and was the last student to defend a dissertation before him.Thell, A., Kärnefelt, I., Seaward, M., & Westberg, M. (Eds.) (2013). In the footsteps of Erik Acharius. 20th biennial meeting of the Nordic Lichen Society. Vadstena 11–15 August 2013. Programme and Abstracts. Nordic Lichen Society. Acharius's dissert ...
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Kurt Polycarp Joachim Sprengel
Kurt Polycarp Joachim Sprengel (3 August 1766 – 15 March 1833) was a German botanist and physician who published an influential multivolume history of medicine, ''Versuch einer pragmatischen Geschichte der Arzneikunde'' (1792–99 in four volumes with later editions running to five) and several other medical reference works. Biography Sprengel was born at Boldekow in Pomerania, and he is considered of German nationality. His father, a clergyman, provided him with a thorough education of wide scope; as boy he distinguished himself as a linguist, in Latin and Greek, and also Arabic; his uncle, Christian Konrad Sprengel (1750–1816), is remembered for his studies in the fertilization of flowers by insects – a subject in which he reached conclusions many years ahead of his time. Spreng. appeared as an author at the age of fourteen, publishing a small work called '' Anleitung zur Botanik für Frauenzimmer'' ("guide to botany for women") in 1780. In 1784 he began to study the ...
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Fruticose Lichen
A fruticose lichen is a form of lichen fungi that is characterized by a coral-like shrubby or bushy growth structure. It is formed from a symbiotic relationship of a photobiont such as green algae or less commonly cyanobacteria and one, two or more mycobionts. Fruticose lichens are not a monophyletic and holophyletic lineage, but is a form encountered in many classes. Fruticose lichens have a complex vegetation structure, and are characterized by an ascending, bushy or pendulous appearance. As with other lichens, many fruticose lichens can endure high degrees of desiccation. They grow slowly and often occur in habitats such as on tree barks, on rock surfaces and on soils in the Arctic and mountain regions. Characteristics Fruticose lichens are lichens composed of a shrubby or bushy thallus and a holdfast. The thallus is the vegetative body of a lichen that does not have true leaves, stems, or roots. The thallus colour is affected by the algae in the lichen, compounds created by t ...
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Cladonia
''Cladonia'' is a genus of moss-like lichens in the family Cladoniaceae. They are the primary food source for reindeer/caribou. ''Cladonia'' species are of economic importance to reindeer-herders, such as the Sami in Scandinavia or the Nenets in Russia. Antibiotic compounds are extracted from some species to create antibiotic cream. The light green species ''Cladonia stellaris'' is used in flower decorations. Although the phylogeny of the genus ''Cladonia'' is still under investigation, two main morphological groups are commonly differentiated by taxonomists: the ''Cladonia'' morpho-type and the ''Cladina'' morpho-type. The ''Cladonia'' morpho-type has many more species, and is generally described as a group of squamulose (grow from squamules), cup-bearing lichens. The ''Cladina'' morpho-types are often referred to as forage lichens, mat-forming lichens, or reindeer lichens (due to their importance as caribou winter forage). ''Cladonia perforata'' ("perforate cladonia") is o ...
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Cladoniaceae
The Cladoniaceae are a family of lichenized fungi in the order Lecanorales. It is one of the largest families of lichen-forming fungi, with about 560 species distributed amongst 17 genera. The reindeer moss and cup lichens (''Cladonia'') belong to this family. The latter genus, which comprises about 500 species, forms a major part of the diet of large mammals in taiga and tundra ecosystems. Many Cladoniaceae lichens grow on soil, but other can use decaying wood, tree trunks, and, in a few instances, rocks as their substrate. They grow in places with high humidity, and cannot tolerate aridity. Many Cladoniaceae species are characterized by a thallus that has two distinct forms: a scaly or crust-like primary thallus that, depending on the species, can be permanent or temporary, and a secondary fruticose thallus called a podetium or pseudopodetium. Cladoniaceae members form symbiotic associations with green algae from the class Trebouxiophyceae, usually the genus '' Asterochloris' ...
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Lichen Species
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

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Taxa Named By Erik Acharius
In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and given a particular ranking, especially if and when it is accepted or becomes established. It is very common, however, for taxonomists to remain at odds over what belongs to a taxon and the criteria used for inclusion. If a taxon is given a formal scientific name, its use is then governed by one of the nomenclature codes specifying which scientific name is correct for a particular grouping. Initial attempts at classifying and ordering organisms (plants and animals) were set forth in Carl Linnaeus's system in '' Systema Naturae'', 10th edition (1758), as well as an unpublished work by Bernard and Antoine Laurent de Jussieu. The idea of a unit-based system of biological classification was first made widely available in 1805 in the i ...
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