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Carpinus Lipoensis
Hornbeams are hardwood trees in the flowering plant genus ''Carpinus'' in the birch family Betulaceae. The 30–40 species occur across much of the temperate regions of the Northern Hemisphere. Origin of names The common English name ''hornbeam'' derives from the hardness of the woods (likened to horn) and the Old English ''beam'' "tree" (cognate with Dutch ‘’Boom’’ and German ''Baum''). The American hornbeam is also occasionally known as blue-beech, ironwood, or musclewood, the first from the resemblance of the bark to that of the American beech ''Fagus grandifolia'', the other two from the hardness of the wood and the muscled appearance of the trunk and limbs. The botanical name for the genus, ''Carpinus'', is the original Latin name for the European species, although some etymologists derive it from the Celtic for a yoke. Taxonomy Formerly some taxonomists segregated them with the genera ''Corylus'' ( hazels) and ''Ostrya'' (hop-hornbeams) in a separate family, Cor ...
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Ypresian
In the geologic timescale the Ypresian is the oldest age (geology), age or lowest stage (stratigraphy), stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by the Eocene Lutetian Age. The Ypresian is consistent with the lower Eocene. Events The Ypresian Age begins during the throes of the Paleocene–Eocene Thermal Maximum (PETM). The Fur Formation in Denmark, the Messel shales in Germany, the Oise amber of France and Cambay amber of India are of this age. The Eocene Okanagan Highlands are an uplands subtropical to temperate series of lakes from the Ypresian. Stratigraphic definition The Ypresian Stage was introduced in scientific literature by Belgium, Belgian geologist André Hubert Dumont in 1850. The Ypresian is named after the Flanders, Flemish city of Ypres in Belgium (spelled ''Ieper'' in Dutch). The definitions of the original stage were totally different from the modern ones. The Ypresi ...
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Hazel
The hazel (''Corylus'') is a genus of deciduous trees and large shrubs native to the temperate Northern Hemisphere. The genus is usually placed in the birch family Betulaceae,Germplasmgobills Information Network''Corylus''Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins .Huxley, A., ed. (1992). ''New RHS Dictionary of Gardening''. Macmillan . though some botanists split the hazels (with the hornbeams and allied genera) into a separate family Corylaceae. The fruit of the hazel is the hazelnut. Hazels have simple, rounded leaves with double-serrate margins. The flowers are produced very early in spring before the leaves, and are monoecious, with single-sex catkins. The male catkins are pale yellow and long, and the female ones are very small and largely concealed in the buds, with only the bright-red, 1-to-3 mm-long styles visible. The fruits are nuts long and 1–2 cm diameter, surrounded by an involucre (husk) which partly to fully encloses the nut. ...
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Catkin
A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind-pollinated (anemophilous) but sometimes insect-pollinated (as in ''Salix''). They contain many, usually unisexual flowers, arranged closely along a central stem that is often drooping. They are found in many plant families, including Betulaceae, Fagaceae, Moraceae, and Salicaceae. Occurrence Catkin-bearing plants include many trees or shrubs such as birch, willow, aspen, hickory, sweet chestnut, and sweetfern (''Comptonia''). In many of these plants, only the male flowers form catkins, and the female flowers are single ( hazel, oak), a cone (alder), or other types (mulberry). In other plants (such as poplar), both male and female flowers are borne in catkins. In Britain, they can be seen in January or February, when many trees are bare for winter. They can even occur in December. Evolution For some time, catkins were believed to be a key synapomorphy among the ...
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Deciduous
In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, after flowering; and to the shedding of ripe fruit. The antonym of ''deciduous'' in the botanical sense is evergreen. Generally, the term "deciduous" means "the dropping of a part that is no longer needed or useful" and the "falling away after its purpose is finished". In plants, it is the result of natural processes. "Deciduous" has a similar meaning when referring to animal parts, such as deciduous antlers in deer, deciduous teeth (baby teeth) in some mammals (including humans); or decidua, the uterine lining that sheds off after birth. Botany In botany and horticulture, deciduous plants, including trees, shrubs and herbaceous perennials, are those that lose all of their leaves for part of the year. This process is called abscissio ...
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Carpinus Betulus 001
Hornbeams are hardwood trees in the flowering plant genus ''Carpinus'' in the birch family Betulaceae. The 30–40 species occur across much of the temperate regions of the Northern Hemisphere. Origin of names The common English name ''hornbeam'' derives from the hardness of the woods (likened to horn) and the Old English ''beam'' "tree" (cognate with Dutch ‘’Boom’’ and German ''Baum''). The American hornbeam is also occasionally known as blue-beech, ironwood, or musclewood, the first from the resemblance of the bark to that of the American beech ''Fagus grandifolia'', the other two from the hardness of the wood and the muscled appearance of the trunk and limbs. The botanical name for the genus, ''Carpinus'', is the original Latin name for the European species, although some etymologists derive it from the Celtic for a yoke. Taxonomy Formerly some taxonomists segregated them with the genera ''Corylus'' (hazels) and ''Ostrya'' (hop-hornbeams) in a separate family, Coryla ...
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Republic, Washington
Republic is a city in Ferry County, Washington, United States. The population was 1,073 at the 2010 census, a 12.5% increase over the 2000 census. It is the county seat of Ferry County. It was the largest mining camp in the Republic Mining District, and home to the "Hot Air Line" railway. Geography Republic is located at (48.648159, −118.734947). According to the United States Census Bureau, the city has a total area of , all of it land. Republic is located near the source of the Sanpoil River in a long graben valley bordered by the Okanagan Highlands to the west and the Kettle mountain range to the east. Curlew Lake, long (Elev: ), provides fishing and boating to summer visitors northeast of Republic. Swan Lake is small mountain lake to the south of Republic and also serves to be a popular lake for local residents and tourists to visit. Republic is surrounded by the Colville National Forest and to the south is the Colville Indian Reservation. History Republic Mining ...
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Klondike Mountain Formation
The Klondike Mountain Formation is an Early Eocene (Ypresian) geological formation located in the northeast central area of Washington state. The formation, named for the type location designated in 1962, Klondike Mountain north of Republic, Washington, is composed of volcanic rocks in the upper unit and volcanics plus lacustrine (lakebed) sedimentation in which a lagerstätte with exceptionally well-preserved plant and insect fossils has been found, along with fossil epithermal hot springs. The formation is the youngest in a group of formations which belong to the Challis Sequence rocks. The formation unconformably overlies rocks of the Eocene Sanpoil Volcanics and much older Triassic and Permian formations. The formation is bounded on its edges by a series of high-angle strike slip faults, which have contained the Klondike Mountain Formation in a series of graben structures, such as the Republic Graben. Public access to a fossiliferous outcrop at the north end of Republic is ...
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Carpinus Perryae
''Carpinus perryae'' is an extinct species of hornbeam known from fossil fruits found in the Klondike Mountain Formation deposits of northern Washington state, dated to the early Eocene Ypresian stage. Based on described features, ''C. perryae'' is the oldest definite species in the genus ''Carpinus''. History and classification ''Carpinus perryae'' was described from two type specimens, the holotype and paratype fertile bracts, numbered UWBM 71171, specimen A and B respectively. Both compression fossil specimens are preserved together on a single rock slab that is part of the Burke Museum paleobotanical collections. Found at the Klondike Mountain Formations UWBM site B2737, which is designated the type locality, the fossils were described by paleobotanists Pigg, Manchester, and Wehr (2003) along with ''Corylus johnsonii'' and '' Palaeocarpinus barksdaleae''. The specific epithet ''perryae'' was coined as a matronym recognizing Madilane Perry for her work founding the Stonero ...
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Early Eocene
In the geologic timescale the Ypresian is the oldest age or lowest stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by the Eocene Lutetian Age. The Ypresian is consistent with the lower Eocene. Events The Ypresian Age begins during the throes of the Paleocene–Eocene Thermal Maximum (PETM). The Fur Formation in Denmark, the Messel shales in Germany, the Oise amber of France and Cambay amber of India are of this age. The Eocene Okanagan Highlands are an uplands subtropical to temperate series of lakes from the Ypresian. Stratigraphic definition The Ypresian Stage was introduced in scientific literature by Belgian geologist André Hubert Dumont in 1850. The Ypresian is named after the Flemish city of Ypres in Belgium (spelled ''Ieper'' in Dutch). The definitions of the original stage were totally different from the modern ones. The Ypresian shares its name with the Belgian Ieper Group (French: ...
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Paraphyletic
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic group (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of Synapomorphy and apomorphy, synapomorphies and symplesiomorphy, symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term was coined by Willi Hennig to apply to well-known taxa like Reptilia (reptiles) which, as commonly named and traditionally defined, is paraphyletic with respect to mammals and birds. Reptilia contains the last common ancestor of reptiles a ...
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Phylogeny
A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological species or other entities based upon similarities and differences in their physical or genetic characteristics. All life on Earth is part of a single phylogenetic tree, indicating common ancestry. In a ''rooted'' phylogenetic tree, each node with descendants represents the inferred most recent common ancestor of those descendants, and the edge lengths in some trees may be interpreted as time estimates. Each node is called a taxonomic unit. Internal nodes are generally called hypothetical taxonomic units, as they cannot be directly observed. Trees are useful in fields of biology such as bioinformatics, systematics, and phylogenetics. ''Unrooted'' trees illustrate only the relatedness of the leaf nodes and do not require the ancestral root to be ...
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Coryloideae
Coryloideae is a subfamily in the woody angiosperm family Betulaceae, commonly known as the birch family, and consists of four extant genera - ''Corylus'' L., ''Ostryopsis'' Decne., ''Carpinus'' L., and ''Ostrya'' Scop. These deciduous trees and shrubs are primarily distributed in the boreal and cool temperate zones of the Northern Hemisphere, with the majority occurring in Asia, many occurring in North America and a few species occurring as far south as South America.Stults, D. Z. & Axsmith, B. J. 2009. Betulaceae From The Pliocene And Pleistocene Of Southwest Alabama, Southeastern United States. ''Review Of Palaeobotany And Palynology'', 155, 25-31. Synapomorphies such reduced staminate flowers, advanced wood anatomy features, and the presence of spermidines in pollen define the Coryloideae.Chen, Z. D., Manchester, S. R. & Sun, H. Y. 1999. Phylogeny And Evolution Of The Betulaceae As Inferred From Dna Sequences, Morphology, And Paleobotany. ''American Journal of Botany'', 86, 1 ...
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