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Canadaspis
''Canadaspis'' ("Shield of Canada") is an extinct genus of bivalved Cambrian arthropod, known from North America and China. They are thought to have been benthic feeders that moved mainly by walking and possibly used its biramous appendages to stir mud in search of food. They have been placed within the Hymenocarina, which includes other bivalved Cambrian arthropods. Description ''Canadaspis'' ''perfecta'' The bivalved carapaces of ''Canadaspis'' ''perfecta'' are typically in length, which taper towards the front end. The head had a small pair of eyes borne on short stalks. Between the eyes is a forward pointing spine, as well as a pair of short antennae, which appear to lack segmentation. Similar antennae are known from '' Waptia'', and are probably homologous to the hemi-ellipsoid bodies of crustaceans, and thus likely have an olfactory function. The head also has another pair of larger, segmented antennae, probably with more than 12 segments, the segments increased in ...
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Canadaspis Laevigata
''Canadaspis'' ("Shield of Canada") is an extinct genus of bivalved Cambrian arthropod, known from North America and China. They are thought to have been benthic feeders that moved mainly by walking and possibly used its biramous appendages to stir mud in search of food. They have been placed within the Hymenocarina, which includes other bivalved Cambrian arthropods. Description ''Canadaspis'' ''perfecta'' The bivalved carapaces of ''Canadaspis'' ''perfecta'' are typically in length, which taper towards the front end. The head had a small pair of eyes borne on short stalks. Between the eyes is a forward pointing spine, as well as a pair of short antennae, which appear to lack segmentation. Similar antennae are known from ''Waptia'', and are probably homologous to the hemi-ellipsoid bodies of crustaceans, and thus likely have an olfactory function. The head also has another pair of larger, segmented antennae, probably with more than 12 segments, the segments increased in lengt ...
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Hymenocarina
Hymenocarina is an order of extinct arthropods known from the Cambrian. They possess bivalved carapaces, typically with exposed posteriors. Members of the group are morphologically diverse and had a variety of ecologies, including as filter feeders and as predators. Recent research has generally considered them to be stem or crown group members of Mandibulata, due the presence of mandibles in at least some species. Taxonomy Hymenocarines are characterized by the combination of following characters: bivalved, convex carapace covering cephalothoracic region; cephalothorax bearing multisegmented antennules and rounded mandibles, alongside post-maxillular limbs with spiny, subdivided basis and endopods with well-developed terminal claws; absence of appendages between antennules and mandibles; median sclerite and lobate protrusions located between compound eyes; posterior tagma (abdomen) with ring-like segments and terminated by a pair of well-developed caudal rami. Based on th ...
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Crustacean
Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group can be treated as a subphylum under the clade Mandibulata. It is now well accepted that the hexapods emerged deep in the Crustacean group, with the completed group referred to as Pancrustacea. Some crustaceans (Remipedia, Cephalocarida, Branchiopoda) are more closely related to insects and the other hexapods than they are to certain other crustaceans. The 67,000 described species range in size from '' Stygotantulus stocki'' at , to the Japanese spider crab with a leg span of up to and a mass of . Like other arthropods, crustaceans have an exoskeleton, which they moult to grow. They are distinguished from other groups of arthropods, such as insects, myriapods and chelicerates, by the possession of biramous (two-parted) limbs, and by th ...
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Waptia
''Waptia fieldensis'' is an extinct species of arthropod from the Middle Cambrian Burgess Shale ''Lagerstätte'' of Canada. It grew to a length of , and had a large bivalved carapace and a segmented body terminating into a pair of tail flaps. It was an active swimmer and likely a predator of soft-bodied prey. It is also one of the oldest animals with direct evidence of brood care. ''Waptia fieldensis'' is the only species classified under the genus ''Waptia''. Other specimens of ''Waptia'' are known from the Spence Shale in Utah. Based on the number of individuals, ''Waptia fieldensis'' is the third most abundant arthropod from the Burgess Shale Formation, with thousands of specimens collected. It was among the first fossils found by the American paleontologist Charles D. Walcott in 1909. He described it in 1912 and named it after two mountains near the discovery site – Wapta Mountain and Mount Field, other specimens Although it bears a remarkable resemblance to modern cr ...
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Extinct
Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and recover may have been lost before this point. Because a species' potential range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxa, where a species presumed extinct abruptly "reappears" (typically in the fossil record) after a period of apparent absence. More than 99% of all species that ever lived on Earth, amounting to over five billion species, are estimated to have died out. It is estimated that there are currently around 8.7 million species of eukaryote globally, and possibly many times more if microorganisms, like bacteria, are included. Notable extinct animal species include non-avian dinosaurs, saber-toothed cats, dodos, m ...
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Exopod
The arthropod leg is a form of jointed appendage of arthropods, usually used for walking. Many of the terms used for arthropod leg segments (called podomeres) are of Latin origin, and may be confused with terms for bones: ''coxa'' (meaning hip, plural ''coxae''), ''trochanter'', ''femur'' (plural ''femora''), ''tibia'' (plural ''tibiae''), ''tarsus'' (plural ''tarsi''), ''ischium'' (plural ''ischia''), ''metatarsus'', ''carpus'', ''dactylus'' (meaning finger), ''patella'' (plural ''patellae''). Homologies of leg segments between groups are difficult to prove and are the source of much argument. Some authors posit up to eleven segments per leg for the most recent common ancestor of extant arthropods but modern arthropods have eight or fewer. It has been argued that the ancestral leg need not have been so complex, and that other events, such as successive loss of function of a ''Hox''-gene, could result in parallel gains of leg segments. In arthropods, each of the leg segments artic ...
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Charles Doolittle Walcott
Charles Doolittle Walcott (March 31, 1850February 9, 1927) was an American paleontologist, administrator of the Smithsonian Institution from 1907 to 1927, and director of the United States Geological Survey.Wonderful Life (book) by Stephen Jay Gould published in 1989, Chapter 4 He is famous for his discovery in 1909 of well-preserved fossils, including some of the oldest soft-part imprints, in the Burgess Shale of British Columbia, Canada. Early life Charles Doolittle Walcott was born on March 31, 1850 in New York Mills, New York. His grandfather, Benjamin S. Walcott, moved from Rhode Island in 1822. His father, also Charles Doolittle Walcott, died when Charles Jr. was only two. Walcott was the youngest of four children. He was interested in nature from an early age, collecting minerals and bird eggs and, eventually, fossils. He attended various schools in the Utica area but left at the age of eighteen without completing high school, the end of his formal education. His intere ...
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Moulting
In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is the manner in which an animal routinely casts off a part of its body (often, but not always, an outer layer or covering), either at specific times of the year, or at specific points in its life cycle. In medieval times it was also known as "mewing" (from the French verb "muer", to moult), a term that lives on in the name of Britain's Royal Mews where the King's hawks used to be kept during moulting time before becoming horse stables after Tudor times. Moulting can involve shedding the Epidermis (skin), epidermis (skin), pelage (hair, feathers, fur, wool), or other external layer. In some groups, other body parts may be shed, for example, the entire exoskeleton in arthropods, including the wings in some insects. Examples In birds In birds, moulting is the periodic replacement of feathers by shedding old feathers while producing new o ...
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Metachronal Rhythm
A metachronal rhythm or metachronal wave refers to wavy movements produced by the sequential action (as opposed to synchronized) of structures such as cilia, segments of worms, or legs. These movements produce the appearance of a travelling wave. A Mexican wave is a large scale example of a metachronal wave. This pattern is found widely in nature such as on the cilia of many aquatic organisms such as ctenophores, molluscs, ciliates as well as on the epithelial surfaces of many body organs. Individual cilia, when part of a metachronal wave being used for protist locomotion, individually beat in a pattern similar to the planar stroke of a flagellum. The difference is that the recovery stroke is at 90 degrees to the power stroke, so that the cilia avoid hitting each other. Metachronal rhythms may be seen in the coordinated movements of the legs of millipedes and other multi-legged land invertebrates, as well as in the coordinated movements of social insects. Such metachronal mo ...
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Mandible (arthropod)
250px, The mandibles of a bull ant The mandible (from la, mandibula or mandĭbŭ-lum, a jaw) of an arthropod is a pair of mouthparts used either for biting or cutting and holding food. Mandibles are often simply called jaws. Mandibles are present in the extant subphyla Myriapoda (millipedes and others), Crustacea and Hexapoda (insects etc.). These groups make up the clade Mandibulata, which is currently believed to be the sister group to the rest of arthropods, the clade Arachnomorpha (Chelicerata and Trilobita). Unlike the chelicerae of arachnids, mandibles can often be used to chew food. Mandibulates also differ by having antennae, and also by having three distinct body regions: head, thorax and abdomen. (The cephalothorax (or prosoma) of chelicerates is a fusion of head and thorax.) Insects Insect mandibles are as diverse in form as their food. For instance, grasshoppers and many other plant-eating insects have sharp-edged mandibles that move side to side. Most ...
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Microbial Mat
A microbial mat is a multi-layered sheet of microorganisms, mainly bacteria and archaea, or bacteria alone. Microbial mats grow at interfaces between different types of material, mostly on submerged or moist surfaces, but a few survive in deserts. A few are found as endosymbionts of animals. Although only a few centimetres thick at most, microbial mats create a wide range of internal chemical environments, and hence generally consist of layers of microorganisms that can feed on or at least tolerate the dominant chemicals at their level and which are usually of closely related species. In moist conditions mats are usually held together by slimy substances secreted by the microorganisms. In many cases some of the bacteria form tangled webs of filaments which make the mat tougher. The best known physical forms are flat mats and stubby pillars called stromatolites, but there are also spherical forms. Microbial mats are the earliest form of life on Earth for which there is good fossi ...
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Derek Briggs
Derek Ernest Gilmor Briggs (born 10 January 1950) is an Irish palaeontologist and taphonomist based at Yale University. Briggs is one of three palaeontologists, along with Harry Blackmore Whittington and Simon Conway Morris, who were key in the reinterpretation of the fossils of the Burgess Shale. He is the Yale University G. Evelyn Hutchinson Professor of Geology and Geophysics, Curator of Invertebrate Paleontology at Yale's Peabody Museum of Natural History, and former Director of the Peabody Museum. Education Briggs was educated at Trinity College Dublin where he graduated with a Bachelor of Arts degree in Geology in 1972. He went on to the University of Cambridge to work under British palaeontologist Harry Blackmore Whittington. He was awarded a PhD in 1976 on ''Arthropods from the Burgess Shale, Middle Cambrian, Canada''. Research and Career While at the University of Cambridge, Briggs worked on the fossils of the Middle Cambrian Burgess Shale of British Columbia alongsi ...
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