Archostemata
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Archostemata
The Archostemata are the smallest suborder of beetles, consisting 45 living species in five families. They are an ancient lineage with a number of primitive characteristics. Antennae may be thread-shaped (filiform) or like a string of beads (moniliform). This suborder also contains the only beetles where both sexes are paedogenic, ''Micromalthus debilis''. Modern archostematan beetles are considered rare, but were more diverse during the Mesozoic. The term "Archostemata" is used more broadly by some authors to include both modern archostematans as well as stem-group beetles like " protocoleopterans", which some modern archostematans closely resemble to due to their plesiomorphic morphology. Genetic research suggests that modern archostematans are a monophyletic group. Some genetic studies have recovered archostematans as the sister group of Myxophaga. Taxonomy There are five extant families. * Family Crowsoniellidae Iablokoff-Khnzorian, 1983 * Family Cupedidae Laporte, 1838 * F ...
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Beetle
Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 described species, is the largest of all orders, constituting almost 40% of described insects and 25% of all known animal life-forms; new species are discovered frequently, with estimates suggesting that there are between 0.9 and 2.1 million total species. Found in almost every habitat except the sea and the polar regions, they interact with their ecosystems in several ways: beetles often feed on plants and fungi, break down animal and plant debris, and eat other invertebrates. Some species are serious agricultural pests, such as the Colorado potato beetle, while others such as Coccinellidae (ladybirds or ladybugs) eat aphids, scale insects, thrips, and other plant-sucking insects that damage crops. Beetles typically have a particularly hard e ...
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Micromalthidae
The telephone-pole beetle (''Micromalthus debilis'') is a beetle native to the eastern United States, and the only living representative of the otherwise extinct family Micromalthidae. They have an unusual lifecycle involving asexually reproducing (parthenogenetic) female larvae, as well as non-functional "ghost adults". Taxonomy Classification of ''M. debilis'' was historically controversial and unsettled. The species, first reported by John Lawrence LeConte in 1878, was long considered one of the Polyphaga, and placed in the Lymexylidae or Telegeusidae, or as a family within the Cantharoidea. However, characteristics of larvae, wings, and male genitalia show that it is in the suborder Archostemata, where it has been placed since 1999. Morphology and life cycle The adult beetle is elongated, ranging from in length, and a dark brown to blackish color, with brownish-yellow legs and antennae. The head is larger than the thorax, with large eyes protruding from either side. ...
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Micromalthus Debilis
The telephone-pole beetle (''Micromalthus debilis'') is a beetle native to the eastern United States, and the only living representative of the otherwise extinct family Micromalthidae. They have an unusual lifecycle involving asexually reproducing (parthenogenetic) female larvae, as well as non-functional "ghost adults". Taxonomy Classification of ''M. debilis'' was historically controversial and unsettled. The species, first reported by John Lawrence LeConte in 1878, was long considered one of the Polyphaga, and placed in the Lymexylidae or Telegeusidae, or as a family within the Cantharoidea. However, characteristics of larvae, wings, and male genitalia show that it is in the suborder Archostemata, where it has been placed since 1999. Morphology and life cycle The adult beetle is elongated, ranging from in length, and a dark brown to blackish color, with brownish-yellow legs and antennae. The head is larger than the thorax, with large eyes protruding from either side. ...
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Telephone-pole Beetle
The telephone-pole beetle (''Micromalthus debilis'') is a beetle native to the eastern United States, and the only living representative of the otherwise extinct family Micromalthidae. They have an unusual lifecycle involving asexually reproducing (parthenogenetic) female larvae, as well as non-functional "ghost adults". Taxonomy Classification of ''M. debilis'' was historically controversial and unsettled. The species, first reported by John Lawrence LeConte in 1878, was long considered one of the Polyphaga, and placed in the Lymexylidae or Telegeusidae, or as a family within the Cantharoidea. However, characteristics of larvae, wings, and male genitalia show that it is in the suborder Archostemata, where it has been placed since 1999. Morphology and life cycle The adult beetle is elongated, ranging from in length, and a dark brown to blackish color, with brownish-yellow legs and antennae. The head is larger than the thorax, with large eyes protruding from either side. ...
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Archostemata
The Archostemata are the smallest suborder of beetles, consisting 45 living species in five families. They are an ancient lineage with a number of primitive characteristics. Antennae may be thread-shaped (filiform) or like a string of beads (moniliform). This suborder also contains the only beetles where both sexes are paedogenic, ''Micromalthus debilis''. Modern archostematan beetles are considered rare, but were more diverse during the Mesozoic. The term "Archostemata" is used more broadly by some authors to include both modern archostematans as well as stem-group beetles like " protocoleopterans", which some modern archostematans closely resemble to due to their plesiomorphic morphology. Genetic research suggests that modern archostematans are a monophyletic group. Some genetic studies have recovered archostematans as the sister group of Myxophaga. Taxonomy There are five extant families. * Family Crowsoniellidae Iablokoff-Khnzorian, 1983 * Family Cupedidae Laporte, 1838 * F ...
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Ommatidae
The Ommatidae are a family of beetles in the suborder Archostemata. The Ommatidae are considered the extant beetle family that has most ancestral characteristics. There are only seven extant species, confined to Australia and South America. However, the geographical distribution was much wider during the Mesozoic spanning across Eurasia and Australia, suggesting that they were widespread on Pangea. So far, over 26 extinct genera containing over 170 species of these beetles have been described. Three extant genera have been assigned to this family: ''Omma,'' '' Tetraphalerus'' and '' Beutelius''. The family is considered to be a subfamily of Cupedidae by some authors, but have been found to be more closely related to Micromalthidae in molecular phylogenies. A close relationship with Micromalthidae is supported by several morphological characters, including those of the mandibles and male genitalia. Due to their rarity, their ecology is obscure, it is likely that their larvae feed ...
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Crowsoniellidae
The Crowsoniellidae are a monotypic family of beetles, in the suborder Archostemata. So far, only a single species, ''Crowsoniella relicta'', has been attributed to this family. Known only from three male specimens collected in 1973 in Italy by Roberto Pace. In a degraded pasture, the beetles were found among the roots of a large hawthorn tree, in deep calcareous soil. No other specimens have been found since. At only in length, this species is comparatively small for this suborder. They also feature several unusual morphological features such as missing alae, modified and reduced mouth parts, and smooth elytra An elytron (; ; , ) is a modified, hardened forewing of beetles (Coleoptera), though a few of the true bugs (Hemiptera) such as the family Schizopteridae are extremely similar; in true bugs, the forewings are called hemelytra (sometimes alterna ... (lacking window punctures characteristic of this suborder). Nothing is known about this species biology. References ...
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Protocoleoptera
The Protocoleoptera are a paraphyletic group of extinct beetles, containing the earliest and most primitive lineages of beetles. They represented the dominant group of beetles during the Permian, but were largely replaced by modern beetle groups during the following Triassic. Protocoleopterans typically possess prognathous (horizontal) heads, distinctive elytra with regular window punctures, culticles with tubercles or scales, as well as a primitive pattern of ventral sclerites, similar to the modern archostematan families Ommatidae and Cupedidae. They are thought to have been xylophagous and wood boring. Kirejtshuk ''et al.'' (2014) argue that the name "Protocoleoptera" should not be used for the group, as Protocoleoptera was originally proposed for the family Protocoleidae , now considered a member of the extinct order Protelytroptera (a stem-group of the modern Dermaptera Earwigs make up the insect order Dermaptera. With about 2,000 species in 12 families, they are ...
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Jurodidae
Jurodidae is a family of beetles that was originally described for the extinct genus '' Jurodes'', known from the Middle-Late Jurassic of Asia. In 1996, a living species, ''Sikhotealinia zhiltzovae'' was discovered in the Sikhote-Alin mountains in Siberia, and assigned to this family. Their placement is uncertain, but are usually considered archostematans. In one study, ''Sikhotealinia'' and ''Jurodes'' were considered a sister group to all other archostematan beetles. However, other authors have considered them to mix characteristics of Archostemata, as well as Polyphaga and Adephaga. Subdivision * '' Jurodes'' Ponomarenko 1985 **''Jurodes ignoramus'' Ponomarenko, 1985 Ichetuy Formation, Russia, Late Jurassic ( Oxfordian) ** ''Jurodes minor'' Ponomarenko, 1990 Glushkovo Formation, Russia, Late Jurassic (Tithonian) ** ''Jurodes daohugouensis'' Dahougou, China, Middle Jurassic (Callovian In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, ...
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Myxophaga
Myxophaga is the second-smallest suborder of the Coleoptera after Archostemata, consisting of roughly 65 species of small to minute beetles in four families. The members of this suborder are aquatic and semiaquatic, and feed on algae. Description Myxophaga have several diagnostic features: the antennae are more or less distinctly clubbed with usually fewer than nine segments, mesocoxal cavities are open laterally and bordered by a mesepimeron and metanepisternum, the hind wings are rolled apically in the resting positions. Internally, they are characterised by the presence of six malpighian tubules and the testes are tube-like and coiled. Beetles of this suborder are adapted to feed on algae. Their mouthparts are characteristic in lacking galeae and having a mobile tooth on their left mandible. Taxonomy There are four extant families in the suborder Myxophaga divided between two superfamilies, containing about 65 described species, and at least one extinct family. Superfami ...
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List Of Subgroups Of The Order Coleoptera
This article scientific classification, classifies the subgroups of the order Coleoptera (beetles) down to the level of family (biology), families, following the system in "Family-group names in Coleoptera (Insecta)", Bouchard, et al. (2011), with corrections and additions from 2020, with common names from bugguide.net. *Order Coleoptera **Suborder †Protocoleoptera ***Superfamily †Tshekardocoleoidea Rohdendorf, 1944 ****Family †Tshekardocoleidae Rohdendorf, 1944 ****Family †Labradorocoleidae Ponomarenko, 1969 ****Family †Oborocoleidae Kukalová, 1969 ***Superfamily †Permocupedoidea Martynov, 1933 ****Family †Permocupedidae Martynov, 1933 ****Family †Taldycupedidae Rohdendorf, 1961 ***Superfamily †Permosynoidea Tillyard, 1924 ****Family †Ademosynidae Ponomarenko, 1968 ****Family †Permosynidae Tillyard, 1924 **Suborder Archostemata ***Superfamily Cupedoidea Laporte, 1836 ****Family Crowsoniellidae Iablokoff-Khnzorian, 1983 ****Family Cupedidae Laporte, 1836 ****F ...
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Tenomerga Mucida
''Tenomerga'' is a genus of beetles in the family Cupedidae. This genus has about 17 extant species, which are native to the eastern Palearctic realm, Palearctic, Nearctic and Indomalayan realm, Oriental regions. Species These 14 species belong to the genus ''Tenomerga'': * ''Tenomerga anguliscutis'' (Kolbe, 1886) * ''Tenomerga cinerea'' (Say, 1831) * ''Tenomerga favella'' Neboiss, 1984 * ''Tenomerga gaolingziensis'' Ge and Yang, 2004 * ''Tenomerga japonica'' (Tamanuki, 1928) * ''Tenomerga kapnodes'' Neboiss, 1984 * ''Tenomerga kurosawai'' Miyatake, 1986 * ''Tenomerga leucophaea'' (Newman, 1839) * ''Tenomerga moultonii'' (Gestro, 1910) * ''Tenomerga mucida'' (Chevrolat, 1844) * ''Tenomerga sibyllae'' (Klapperich, 1950) * ''Tenomerga tianmuensis'' Ge and Yang, 2004 * ''Tenomerga trabecula'' Neboiss, 1984 * ''Tenomerga yamato'' Miyatake, 1985 i c g Data sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net References External links

* * Archostemata g ...
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