Blumeria Graminis F.sp. Tritici Resistance
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Blumeria Graminis F.sp. Tritici Resistance
''Blumeria graminis'' (commonly called barley powdery mildew or corn mildew) is a fungus that causes powdery mildew on grasses, including cereals. It is the only species in the genus ''Blumeria''. It has also been called ''Erysiphe graminis'' and (by its anamorph) ''Oidium monilioides'' or ''Oidium tritici''. Systematics Previously ''B. graminis'' was included within the genus ''Erysiphe'', but molecular studies have placed it into a clade of its own. Thus since 1975, the species ''graminis'' was moved into the new taxa ''Blumeria'' of which it is the only species. ''Blumeria'' differs from ''Erysiphe'' in its digitate haustoria and in details of the conidial wall. As well ''Blumeria'' is considered to be phylogenetically distinct from ''Erisiphe'' as it is a plant pathogen that hosts solely on the true grasses of ''Poaceae. '' Eight special forms or ''formae speciales'' (ff.spp.) of ''B. graminis'' have been distinguished, each of which is parasitic on a particular genus of gras ...
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Fungi
A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from the other eukaryotic kingdoms, which by one traditional classification include Plantae, Animalia, Protozoa, and Chromista. A characteristic that places fungi in a different kingdom from plants, bacteria, and some protists is chitin in their cell walls. Fungi, like animals, are heterotrophs; they acquire their food by absorbing dissolved molecules, typically by secreting digestive enzymes into their environment. Fungi do not photosynthesize. Growth is their means of mobility, except for spores (a few of which are flagellated), which may travel through the air or water. Fungi are the principal decomposers in ecological systems. These and other differences place fungi in a single group of related organisms, named the ''Eumycota'' (''t ...
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Wheat
Wheat is a grass widely cultivated for its seed, a cereal grain that is a worldwide staple food. The many species of wheat together make up the genus ''Triticum'' ; the most widely grown is common wheat (''T. aestivum''). The archaeological record suggests that wheat was first cultivated in the regions of the Fertile Crescent around 9600 BCE. Botanically, the wheat kernel is a type of fruit called a caryopsis. Wheat is grown on more land area than any other food crop (, 2014). World trade in wheat is greater than for all other crops combined. In 2020, world production of wheat was , making it the second most-produced cereal after maize. Since 1960, world production of wheat and other grain crops has tripled and is expected to grow further through the middle of the 21st century. Global demand for wheat is increasing due to the unique viscoelastic and adhesive properties of gluten proteins, which facilitate the production of processed foods, whose consumption is inc ...
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Ascospore
An ascus (; ) is the sexual spore-bearing cell produced in ascomycete fungi. Each ascus usually contains eight ascospores (or octad), produced by meiosis followed, in most species, by a mitotic cell division. However, asci in some genera or species can occur in numbers of one (e.g. ''Monosporascus cannonballus''), two, four, or multiples of four. In a few cases, the ascospores can bud off conidia that may fill the asci (e.g. ''Tympanis'') with hundreds of conidia, or the ascospores may fragment, e.g. some ''Cordyceps'', also filling the asci with smaller cells. Ascospores are nonmotile, usually single celled, but not infrequently may be coenocytic (lacking a septum), and in some cases coenocytic in multiple planes. Mitotic divisions within the developing spores populate each resulting cell in septate ascospores with nuclei. The term ocular chamber, or oculus, refers to the epiplasm (the portion of cytoplasm not used in ascospore formation) that is surrounded by the "bourrelet ...
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Ascus
An ascus (; ) is the sexual spore-bearing cell produced in ascomycete fungi. Each ascus usually contains eight ascospores (or octad), produced by meiosis followed, in most species, by a mitotic cell division. However, asci in some genera or species can occur in numbers of one (e.g. ''Monosporascus cannonballus''), two, four, or multiples of four. In a few cases, the ascospores can bud off conidia that may fill the asci (e.g. ''Tympanis'') with hundreds of conidia, or the ascospores may fragment, e.g. some ''Cordyceps'', also filling the asci with smaller cells. Ascospores are nonmotile, usually single celled, but not infrequently may be coenocytic (lacking a septum), and in some cases coenocytic in multiple planes. Mitotic divisions within the developing spores populate each resulting cell in septate ascospores with nuclei. The term ocular chamber, or oculus, refers to the epiplasm (the portion of cytoplasm not used in ascospore formation) that is surrounded by the "bourrelet ...
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Ascocarp
An ascocarp, or ascoma (), is the fruiting body ( sporocarp) of an ascomycete phylum fungus. It consists of very tightly interwoven hyphae and millions of embedded asci, each of which typically contains four to eight ascospores. Ascocarps are most commonly bowl-shaped (apothecia) but may take on a spherical or flask-like form that has a pore opening to release spores (perithecia) or no opening (cleistothecia). Classification The ascocarp is classified according to its placement (in ways not fundamental to the basic taxonomy). It is called ''epigeous'' if it grows above ground, as with the morels, while underground ascocarps, such as truffles, are termed ''hypogeous''. The structure enclosing the hymenium is divided into the types described below (apothecium, cleistothecium, etc.) and this character ''is'' important for the taxonomic classification of the fungus. Apothecia can be relatively large and fleshy, whereas the others are microscopic—about the size of flecks ...
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Mycelium
Mycelium (plural mycelia) is a root-like structure of a fungus consisting of a mass of branching, thread-like hyphae. Fungal colonies composed of mycelium are found in and on soil and many other substrate (biology), substrates. A typical single spore germinates into a Monokaryon, monokaryotic mycelium, which cannot reproduce sexually; when two compatible monokaryotic mycelia join and form a dikaryotic mycelium, that mycelium may form sporocarp (fungi), fruiting bodies such as mushrooms. A mycelium may be minute, forming a colony that is too small to see, or may grow to span thousands of acres as in ''Armillaria''. Through the mycelium, a fungus absorbs nutrients from its environment. It does this in a two-stage process. First, the hyphae secrete enzymes onto or into the food source, which break down biopolymers, biological polymers into smaller units such as monomers. These monomers are then absorbed into the mycelium by facilitated diffusion and active transport. Mycelia are v ...
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Lolium
''Lolium'' is a genus of tufted grasses in the bluegrass subfamily (Pooideae). It is often called ryegrass, but this term is sometimes used to refer to grasses in other genera. They are characterized by bunch-like growth habits. ''Lolium'' is native to Europe, Asia and northern Africa, as well as being cultivated and naturalized in Australia, the Americas, and various oceanic islands. Ryegrasses are naturally diploid, with 2n=14, and are closely related to the fescues (''Festuca''). Ryegrass should not be confused with rye, which is a grain crop. Species Species of ''Lolium'' include: * ''Lolium arundinaceum'' (Schreb.) Darbysh. - Eurasia + North Africa from Portugal + Canary Islands to Himalayas + Xinjiang; naturalized in East Asia, Australia, North + South America, various islands * ''Lolium canariense'' Steud. - Canary Islands ryegrass - Canary Islands, Cape Verde * ''Lolium giganteum '' Lam. - Eurasia from Ireland to China; Bioko * ''Lolium hybridum'' Hausskn. - Assam, B ...
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Bromus
''Bromus'' is a large genus of grasses, classified in its own tribe Bromeae. They are commonly known as bromes, brome grasses, cheat grasses or chess grasses. Estimates in the scientific literature of the number of species have ranged from 100 to 400, but plant taxonomists currently recognize around 160–170 species. ''Bromus'' is part of the cool-season grass lineage (subfamily Pooideae), which includes about 3300 species. Within Pooideae, ''Bromus'' is classified in tribe Bromeae (it is the only genus in the tribe). ''Bromus'' is closely related to the wheat-grass lineage (tribe Triticeae) that includes such economically important genera as ''Triticum'' (wheat), ''Hordeum'' (barley) and '' Secale'' (rye). Etymology The generic name ''Bromus'' is derived from the Latin ''bromos'', a borrowed word from the Ancient Greek (). and mean ''oats'', but seems to have referred specifically to ''Avena sativa'' (Hippocrates ''On Regimen in Acute Diseases'' 2.43, Dioscorides Med ...
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Elymus (plant)
''Elymus'' is a genus of perennial plants with approximately 150 species in the grass family, related to rye, wheat, and other widely grown cereal grains. ''Elymus'' is a cosmopolitan genus, represented by species across all continents of the world. Common names include couch grass, wildrye and wheatgrass. Species Species accepted by the Plants of the World Online as of 2021: ref name=POWO> *''Elymus abolinii'' *''Elymus aenaeanus'' *'' Elymus afghanicus'' *'' Elymus africanus'' *'' Elymus alaskanus'' *'' Elymus albicans'' *'' Elymus alienus'' *'' Elymus alpinus'' *'' Elymus altissimus'' *'' Elymus amgensis'' *'' Elymus angsaiensis'' *''Elymus angulatus'' *'' Elymus angustispiculatus'' *''Elymus anthosachnoides'' *'' Elymus antiquus'' *'' Elymus × apiculatus'' *''Elymus arcuatus'' *''Elymus aristiglumis'' *''Elymus arizonicus'' *''Elymus athericus'' *''Elymus atratus'' *''Elymus bakeri'' *''Elymus barbicallus'' *''Elymus barystachyus'' *''Elymus bori ...
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Agropyron
''Agropyron'' is a genus of Eurasian plants in the grass family), native to Europe and Asia but widely naturalized in North America. Species in the genus are commonly referred to as wheatgrass. ; Species * ''Agropyron badamense'' - Tajikistan, Kyrgyzstan, Uzbekistan, Kazakhstan * ''Agropyron bulbosum'' - Iran * ''Agropyron cimmericum'' - Ukraine, Crimea * ''Agropyron cristatum'' - Crested wheatgrass - Eurasia + North Africa from Spain + Morocco to Korea + Khabarovsk; naturalized in western + central North America (United States, Canada, northern Mexico) * ''Agropyron dasyanthum'' - Ukraine * ''Agropyron desertorum'' - Desert Wheatgrass - from Crimea + Caucasus to Mongolia + Siberia * ''Agropyron deweyi'' - Turkey * ''Agropyron fragile'' - Siberian wheatgrass - from Caucasus to Mongolia; naturalized in scattered locales in western United States + Canada * ''Agropyron michnoi'' - Buryatiya, Zabaykalsky Krai, Mongolia, Inner Mongolia * ''Agropyron mongolicum'' - Gansu, Inner ...
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Oats
The oat (''Avena sativa''), sometimes called the common oat, is a species of cereal grain grown for its seed, which is known by the same name (usually in the plural, unlike other cereals and pseudocereals). While oats are suitable for human consumption as oatmeal and rolled oats, one of the most common uses is as livestock feed. Oats are a nutrient-rich food associated with lower blood cholesterol when consumed regularly. Avenins are oat gluten proteins, similar to gliadin in wheat. They can trigger celiac disease in a small proportion of people. Also, oat products are frequently contaminated by other gluten-containing grains, mainly wheat and barley. Origin The wild ancestor of ''Avena sativa'' and the closely related minor crop '' A. byzantina'' is '' A. sterilis''. ''A. sterilis'' is a wild oat that is naturally hexaploid. Genetic evidence shows the ancestral forms of ''A. sterilis'' grew in the Fertile Crescent of the Near East. Oats are usually thought to have emerged a ...
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