Benthosuchidae
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Benthosuchidae
''Benthosuchus'' (meaning "deep water crocodile") is an extinct genus of temnospondyl amphibian from the Early Triassic of Russia. It was primarily aquatic, living in rivers and lakes. Multiple species are known, with the largest reaching about 2.5 meters in length. Russian paleontologist Ivan Yefremov fremovcalled the genus ''Benthosaurus'' "deep water lizard" (from Ancient Greek βένθος (''benthos'') "depth, deep water") in his original 1929 description, "in view of its clearly indicated adaptation to life in deep water" shown by "the position of the orbits and the flatness of the skull." The type species ''B. sushkini'' honored his late teacher Petr Sushkin. The generic name was preoccupied by ''Benthosaurus'' Goode & Bean, 1886, a fish, and he renamed the genus ''Benthosuchus'' ("deep water crocodile") in 1937.Efremov, I. A. (1929). ''Benthosaurus sushkini'', ein neuer Labyrinthodont der permotriassischen Ablagerungen der Sharschenga Flusses. ''Bulletin of the Academ ...
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Benthosuchus2DB2small
''Benthosuchus'' (meaning "deep water crocodile") is an extinct genus of temnospondyli, temnospondyl amphibian from the Early Triassic of Russia. It was primarily aquatic, living in rivers and lakes. Multiple species are known, with the largest reaching about 2.5 meters in length. Russian paleontologist Ivan Yefremov [Efremov] called the genus ''Benthosaurus'' "deep water lizard" (from Ancient Greek βένθος (''benthos'') "depth, deep water") in his original 1929 description, "in view of its clearly indicated adaptation to life in deep water" shown by "the position of the orbits and the flatness of the skull." The type species ''B. sushkini'' honored his late teacher Petr Sushkin. The generic name was preoccupied by ''Benthosaurus'' Goode & Bean, 1886, a fish, and he renamed the genus ''Benthosuchus'' ("deep water crocodile") in 1937.Efremov, I. A. (1929). ''Benthosaurus sushkini'', ein neuer Labyrinthodont der permotriassischen Ablagerungen der Sharschenga Flusses. ''Bull ...
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Benthosuchus Korobkovi
''Benthosuchus'' (meaning "deep water crocodile") is an extinct genus of temnospondyl amphibian from the Early Triassic of Russia. It was primarily aquatic, living in rivers and lakes. Multiple species are known, with the largest reaching about 2.5 meters in length. Russian paleontologist Ivan Yefremov fremovcalled the genus ''Benthosaurus'' "deep water lizard" (from Ancient Greek βένθος (''benthos'') "depth, deep water") in his original 1929 description, "in view of its clearly indicated adaptation to life in deep water" shown by "the position of the orbits and the flatness of the skull." The type species ''B. sushkini'' honored his late teacher Petr Sushkin. The generic name was preoccupied by ''Benthosaurus'' Goode & Bean, 1886, a fish, and he renamed the genus ''Benthosuchus'' ("deep water crocodile") in 1937.Efremov, I. A. (1929). ''Benthosaurus sushkini'', ein neuer Labyrinthodont der permotriassischen Ablagerungen der Sharschenga Flusses. ''Bulletin of the Academ ...
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Thoosuchus
''Thoosuchus'' (meaning "active crocodile") is an extinct genus of basal trematosauroid trematosaurian temnospondyl. Fossils have been found from Russia and date back to the Early Triassic. It is the type genus of the family Thoosuchidae, formerly called the subfamily Thoosuchinae and placed within Benthosuchidae. Efremov, I. A. (1940). Preliminary description of the new Permian and Triassic Tetrapoda from U.S.S.R. ''Trudy Paleontologicheskogo Instituta'' 10:1–140. The benthosuchids were originally composed of the majority of basal trematosaurian forms regarded as the ancestors of the trematosaurids. Although the genus was first named in 1940, material from one species, ''E. yakovlevi'', was originally tentatively referred to ''Trematosuchus'' in 1926. For its family, ''Thoosuchus'' was fairly small, reaching a little over 60 cm with a 15 cm skull. ''Thoosuchus'' superficially resembles the more derived trematosaurids, but can be distinguished from them on the ...
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Early Triassic
The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which is a unit in chronostratigraphy. The Early Triassic is the oldest epoch of the Mesozoic Era. It is preceded by the Lopingian Epoch (late Permian, Paleozoic Era) and followed by the Middle Triassic Epoch. The Early Triassic is divided into the Induan and Olenekian ages. The Induan is subdivided into the Griesbachian and Dienerian subages and the Olenekian is subdivided into the Smithian and Spathian subages. The Lower Triassic series is coeval with the Scythian Stage, which is today not included in the official timescales but can be found in older literature. In Europe, most of the Lower Triassic is composed of Buntsandstein, a lithostratigraphic unit of continental red beds. The Early Triassic and partly also the Middle Triassic span the in ...
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Capitosauria
Capitosauria is an extinct group of large temnospondyl amphibians with simplified stereospondyl vertebrae. Mainly living as piscivores in lakes and rivers, the Capitosauria and its sister taxon Trematosauria were the only major labyrinthodonts that existed during the Mesozoic in ecological niches broadly similar to those of modern crocodiles, and some grew to very large sizes. At 6 meters in length, the Mid-Triassic '' Mastodonsaurus giganteus'' is not only thought to have been the largest capitosaur, but possibly also the largest amphibian to have lived. The latest known remains are from the Rhaetian of Germany and are referred to ''Cyclotosaurus''. Capitosauria was first named by Schoch and Milner (2000) and further described by Yates and Warren (2000), who assigned ''Lydekkerina'' and Mastodonsauroidea to it. It was described by Damiani (2001) under the name Mastodonsauroidea. In their phylogenetic analysis of temnospondyls, Ruta ''et al.'' (2007) placed ''Lydekkerina'' and it ...
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East Russian Depression
East or Orient is one of the four cardinal directions or points of the compass. It is the opposite direction from west and is the direction from which the Sun rises on the Earth. Etymology As in other languages, the word is formed from the fact that east is the direction where the Sun rises: ''east'' comes from Middle English ''est'', from Old English ''ēast'', which itself comes from the Proto-Germanic *''aus-to-'' or *''austra-'' "east, toward the sunrise", from Proto-Indo-European *aus- "to shine," or "dawn", cognate with Old High German ''*ōstar'' "to the east", Latin ''aurora'' 'dawn', and Greek ''ēōs'' 'dawn, east'. Examples of the same formation in other languages include Latin oriens 'east, sunrise' from orior 'to rise, to originate', Greek ανατολή anatolé 'east' from ἀνατέλλω 'to rise' and Hebrew מִזְרָח mizraḥ 'east' from זָרַח zaraḥ 'to rise, to shine'. ''Ēostre'', a Germanic goddess of dawn, might have been a personificatio ...
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Capitosaurid
Mastodonsauridae is a family of capitosauroid temnospondyls. Fossils belonging to this family have been found in North America, Greenland, Europe, Asia, and Australia. The family Capitosauridae is synonymous with Mastodonsauridae. Description Size Mastodonsaurids were generally large amphibians, with some length estimates ranging from 3 to 6 meters. Some genera, like ''Mastodonsaurus'' and ''Cyclotosaurus'' were specially large, reaching at least 4 meters or more. Other genera like ''Parotosuchus'' and ''Paracyclotosaurus'' only reached 2 meters or more in length, but it's still pretty large compared to modern-day amphibians. Distinguishing features Some mastodonsaurids including ''Parotosuchus'' were covered in a scaly skin, unlike the smooth skin of modern-day amphibians, and probably moved with an eel-like motion in the water. Another peculiar mastodonsaurid is ''Sclerothorax'', with unusual features including its elongated neural spines. The neural spines are tallest at the ...
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Wetlugasaurus
''Wetlugasaurus'' (meaning "Vetluga River lizard") is an extinct genus of Temnospondyli, temnospondyl amphibian from the Early Triassic (Olenekian) Charkabozh Formation, Charkabozh, Kzylsaiskaya Formation, Kzylsaiskaya, Petropavlovka Formation, Petropavlovka, Kamennyi Yar Formation, Kamennyi Yar and Vetluga Series Formations of northern Russia and Greenland. It had a long skull, and reached a total length of . References Further reading

* ''The Age of Dinosaurs in Russia and Mongolia'' by Michael J. Benton, Mikhail A. Shishkin, David M. Unwin, and Evgenii N. Kurochkin. p. 35-59. * ''Bibliography Of Fossil Vertebrates 1934-1938'' by C. I. Camp Capitosaurs Prehistoric amphibian genera Triassic temnospondyls of Europe Olenekian life Fossils of Russia Fossil taxa described in 1930 {{triassic-animal-stub ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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Trematosauridae
Trematosauridae are a family of large marine temnospondyl amphibians with many members. They first appeared during the Induan age of the Early Triassic, and existed until around the Carnian stage of the Late Triassic, although by then they were very rare. By the Middle Triassic they had become widespread throughout Laurasia and Gondwana with fossils being found in Europe, Asia, Madagascar, and Australia. They are one of the most derived families of the Trematosauroidea superfamily in that they are the only family that have fully marine lifestyles. Long, slender snouts that are characteristic of the trematosaurids, with some members having rostrums resembling those of modern-day gavials. Traditionally, two subfamilies within Trematosauridae can be identified, the relatively short-nosed Trematosaurinae and the long-nosed Lonchorhynchinae._A_third_subfamily,_Tertreminae.html" ;"title=".... A third subfamily, Tertreminae">.... A third subfamily, Tertreminae, was named in 2000 ...
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Trematosauria
Trematosauria is one of two major groups of temnospondyl amphibians that survived the Permian-Triassic extinction event, the other (according to Yates and Warren 2000) being the Capitosauria. The trematosaurs were a diverse and important group that included many medium-sized to large forms that were semi-aquatic to totally aquatic. The group included long-snouted forms such as the Trematosauroidea, trematosauroids and short, broad-headed forms such as the Metoposauridae, metoposaurs.Brusatte, S. L., Butler R. J., Mateus O., & Steyer S. J. (2015). A new species of Metoposaurus from the Late Triassic of Portugal and comments on the systematics and biogeography of metoposaurid temnospondyls. Journal of Vertebrate Paleontology. e912988., 2015: Although most groups did not survive beyond the Triassic, one lineage, the Brachyopoidea, brachyopoids, continued until the Cretaceous period. Trematosauria is defined as all Stereospondyli, stereospondyls more closely related to ''Trematosauru ...
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Monotypic
In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispecific" or "monospecific" is sometimes preferred. In botanical nomenclature, a monotypic genus is a genus in the special case where a genus and a single species are simultaneously described. In contrast, an oligotypic taxon contains more than one but only a very few subordinate taxa. Examples Just as the term ''monotypic'' is used to describe a taxon including only one subdivision, the contained taxon can also be referred to as monotypic within the higher-level taxon, e.g. a genus monotypic within a family. Some examples of monotypic groups are: Plants * In the order Amborellales, there is only one family, Amborellaceae and there is only one genus, '' Amborella'', and in this genus there is only one species, namely ''Amborella trichopoda. ...
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