Bellatoripes Fredlundi
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Bellatoripes Fredlundi
''Bellatoripes'' (Latin for "warlike foot") is an ichnogenus of footprint produced by a large theropod dinosaur so far known only from the Late Cretaceous of Alberta and British Columbia in Canada. The tracks are large and three-toed, and based on their size are believed to have been made by tyrannosaurids, such as ''Albertosaurus'' and ''Daspletosaurus''. Fossils of ''Bellatoripes'' are notable for preserving trackways of multiple individual tyrannosaurids all travelling in the same direction at similar speeds, suggesting the prints may have been made by a group, or pack, of tyrannosaurids moving together. Such inferences of behaviour cannot be made with fossil bones alone, so the record of ''Bellatoripes'' tracks together is important for understanding how large predatory theropods such as tyrannosaurids may have lived. Discovery and classification ''Bellatoripes'' was named in 2014 from a tracksite discovered in the Wapiti Formation, British Columbia containing the trackways o ...
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Upper Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and Anta ...
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Ichnospecies
An ichnotaxon (plural ichnotaxa) is "a taxon based on the fossilized work of an organism", i.e. the non-human equivalent of an artifact. ''Ichnotaxa'' comes from the Greek ίχνος, ''ichnos'' meaning ''track'' and ταξις, ''taxis'' meaning ''ordering''.Definition o'ichno'at dictionary.com. Ichnotaxa are names used to identify and distinguish morphologically distinctive ichnofossils, more commonly known as trace fossils. They are assigned genus and species ranks by ichnologists, much like organisms in Linnaean taxonomy. These are known as ichnogenera and ichnospecies, respectively. "Ichnogenus" and "ichnospecies" are commonly abbreviated as "igen." and "isp.". The binomial names of ichnospecies and their genera are to be written in italics. Most researchers classify trace fossils only as far as the ichnogenus rank, based upon trace fossils that resemble each other in morphology but have subtle differences. Some authors have constructed detailed hierarchies up to ichnosupe ...
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Pes (anatomy)
The pes (Latin for ''foot'') is the zoological term for the distal portion of the hind limb of tetrapod animals. It is the part of the pentadactyl limb that includes the metatarsals and digits (phalanges). During evolution, it has taken many forms and served a variety of functions. It can be represented by the foot of primates, the lower hind limb of hoofed animals, the lower portion of the leg of dinosaurs including birds or the rear paw. It is also represented in the rear 'paddle' of extinct marine reptiles, such as plesiosaurs. The oldest types of tetrapods had seven or eight digits. See also *Manus (anatomy) The manus (Latin for ''hand'', plural manus) is the zoological term for the distal portion of the fore limb of an animal. In tetrapods, it is the part of the pentadactyl limb that includes the metacarpals and digits (phalanges). During evolution ... References Vertebrate anatomy Dinosaur anatomy Lower limb anatomy {{Vertebrate anatomy-stub ...
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Tridactyl
In biology, dactyly is the arrangement of digits (fingers and toes) on the hands, feet, or sometimes wings of a tetrapod animal. It comes from the Greek word δακτυλος (''dáktylos'') = "finger". Sometimes the ending "-dactylia" is used. The derived adjectives end with "-dactyl" or "-dactylous". As a normal feature Pentadactyly Pentadactyly (from Greek "five") is the condition of having five digits on each limb. It is traditionally believed that all living tetrapods are descended from an ancestor with a pentadactyl limb, although many species have now lost or transformed some or all of their digits by the process of evolution. However, this viewpoint was challenged by Stephen Jay Gould in his 1991 essay "Eight (Or Fewer) Little Piggies", where he pointed out polydactyly in early tetrapods and described the specializations of digit reduction. Despite the individual variations listed below, the relationship is to the original five-digit model. In reptiles, the limbs ...
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Bellatoripes Photogrammetry
''Bellatoripes'' (Latin for "warlike foot") is an ichnogenus of footprint produced by a large theropod dinosaur so far known only from the Late Cretaceous of Alberta and British Columbia in Canada. The tracks are large and three-toed, and based on their size are believed to have been made by tyrannosaurids, such as ''Albertosaurus'' and ''Daspletosaurus''. Fossils of ''Bellatoripes'' are notable for preserving trackways of multiple individual tyrannosaurids all travelling in the same direction at similar speeds, suggesting the prints may have been made by a group, or pack, of tyrannosaurids moving together. Such inferences of behaviour cannot be made with fossil bones alone, so the record of ''Bellatoripes'' tracks together is important for understanding how large predatory theropods such as tyrannosaurids may have lived. Discovery and classification ''Bellatoripes'' was named in 2014 from a tracksite discovered in the Wapiti Formation, British Columbia containing the trackways o ...
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Synapomorphies
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis T ...
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Stratigraphic
Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithostratigraphy (lithologic stratigraphy), biostratigraphy (biologic stratigraphy), and chronostratigraphy (stratigraphy by age). Historical development Catholic priest Nicholas Steno established the theoretical basis for stratigraphy when he introduced the law of superposition, the principle of original horizontality and the principle of lateral continuity in a 1669 work on the fossilization of organic remains in layers of sediment. The first practical large-scale application of stratigraphy was by William Smith in the 1790s and early 19th century. Known as the "Father of English geology", Smith recognized the significance of strata or rock layering and the importance of fossil markers for correlating strata; he created the first geologic map o ...
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Family (biology)
Family ( la, familia, plural ') is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as the "walnut family". What belongs to a family—or if a described family should be recognized at all—are proposed and determined by practicing taxonomists. There are no hard rules for describing or recognizing a family, but in plants, they can be characterized on the basis of both vegetative and reproductive features of plant species. Taxonomists often take different positions about descriptions, and there may be no broad consensus across the scientific community for some time. The publishing of new data and opini ...
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Mongolia
Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, making it the world's most sparsely populated sovereign nation. Mongolia is the world's largest landlocked country that does not border a closed sea, and much of its area is covered by grassy steppe, with mountains to the north and west and the Gobi Desert to the south. Ulaanbaatar, the capital and largest city, is home to roughly half of the country's population. The territory of modern-day Mongolia has been ruled by various nomadic empires, including the Xiongnu, the Xianbei, the Rouran, the First Turkic Khaganate, and others. In 1206, Genghis Khan founded the Mongol Empire, which became the largest contiguous land empire in history. His grandson Kublai Khan conquered China proper and established the Yuan dynasty. After the co ...
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Nemegt Formation
The Nemegt Formation (also known as Nemegtskaya Svita) is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilians, and a diverse fauna of dinosaurs, including birds. Description The Nemegt Formation is composed of mudstones and sandstones that were deposited by ancient lakes, streams, and flood plains. The Altan Uul locality was described by Michael Novacek as "a canyon carved out of a very rich series of sedimentary rocks" with "steep cliffs and narrow washes". The climate associated with it was wetter than when preceding formations were deposited; there seems to have existed at least some degree of forest cover. Fossilized trunks have been also found. These petrified wood, and the remains of Araucariaceae conifers indicate that the forests of the Nemegt were thickly wooded, with a high canopy formed by tall conifer trees. When examined, the rock ...
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Hallux
Toes are the digits (fingers) of the foot of a tetrapod. Animal species such as cats that walk on their toes are described as being '' digitigrade''. Humans, and other animals that walk on the soles of their feet, are described as being '' plantigrade''; ''unguligrade'' animals are those that walk on hooves at the tips of their toes. Structure There are normally five toes present on each human foot. Each toe consists of three phalanx bones, the proximal, middle, and distal, with the exception of the big toe ( la, hallux). For a minority of people, the little toe also is missing a middle bone. The hallux only contains two phalanx bones, the proximal and distal. The joints between each phalanx are the interphalangeal joints. The proximal phalanx bone of each toe articulates with the metatarsal bone of the foot at the metatarsophalangeal joint. Each toe is surrounded by skin, and present on all five toes is a toenail. The toes are, from medial to lateral: * the first ...
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Tyrannosaurus
''Tyrannosaurus'' is a genus of large theropoda, theropod dinosaur. The species ''Tyrannosaurus rex'' (''rex'' meaning "king" in Latin), often called ''T. rex'' or colloquially ''T-Rex'', is one of the best represented theropods. ''Tyrannosaurus'' lived throughout what is now western North America, on what was then an island continent known as Laramidia. ''Tyrannosaurus'' had a much wider range than other Tyrannosauridae, tyrannosaurids. Fossils are found in a variety of geologic formation, rock formations dating to the Maastrichtian Age (geology), age of the Upper Cretaceous Period (geology), period, 68 to 66 mya (unit), million years ago. It was the last known member of the tyrannosaurids and among the last non-aves, avian dinosaurs to exist before the Cretaceous–Paleogene extinction event. Like other tyrannosaurids, ''Tyrannosaurus'' was a bipedal carnivore with a massive skull balanced by a long, heavy tail. Relative to its large and powerful hind limbs, the foreli ...
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