Batrachognathus
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Batrachognathus
''Batrachognathus'' is an extinct genus of anurognathid pterosaur from the Late Jurassic ( Oxfordian to Kimmeridgian) Karabastau Formation of the central Asian republic of Kazakhstan. The genus was named in 1948 by the Russian paleontologist Anatoly Nicolaevich Ryabinin. The type species is ''Batrachognathus volans''. The genus name is derived from Greek ''batrakhos'', "frog" and ''gnathos'', "jaw", in reference to the short wide head. The specific epithet means "flying" in Latin. Description ''Batrachognathus'' was a small pterosaur, with a wingspan of and body mass of . Like all anurognathids ''Batrachognathus'' is assumed to have been an insectivore, catching insects and perhaps small fish on the wing with its broad mouth. Three fossils have been found in a lacustrine sediment in the North-West Tien Shan foothills of the Karatau Mountains. In the Jurassic this area had some similarities in habitat to the Solnhofen lagoon deposits in Bavaria, Germany. The genus is based on h ...
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Anurognathid Jaws
Anurognathidae is a family of small, short-tailed pterosaurs that lived in Europe, Asia, and possibly North America during the Jurassic and Cretaceous periods. Five genera are known: ''Anurognathus'', from the Late Jurassic of Germany; ''Jeholopterus'', from the Middle to Late Jurassic of China; ''Dendrorhynchoides'', from the Middle Jurassic of China; ''Batrachognathus'', from the Late Jurassic of Kazakhstan; and ''Vesperopterylus'', from the Early Cretaceous of China. Bennett (2007) suggested that the holotype of ''Mesadactylus'', BYU 2024, a synsacrum, belonged to an anurognathid, though this affinity has been questioned by other authors. ''Mesadactylus'' is from the Late Jurassic Morrison Formation of the United States. Indeterminate anurognathid remains have also been reported from the Middle Jurassic Bakhar Svita of Mongolia and the Early Cretaceous of North Korea. Classification A family Anurognathidae was named in 1928 by Franz Nopcsa von Felső-Szilvás (as the subfami ...
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Anurognathidae
Anurognathidae is a family (biology), family of small, short-tailed pterosaurs that lived in Europe, Asia, and possibly North America during the Jurassic and Cretaceous periods. Five genera are known: ''Anurognathus'', from the Late Jurassic of Germany; ''Jeholopterus'', from the Middle Jurassic, Middle to Late Jurassic of China; ''Dendrorhynchoides'', from the Middle Jurassic of China; ''Batrachognathus'', from the Late Jurassic of Kazakhstan; and ''Vesperopterylus'', from the Early Cretaceous of China. Bennett (2007) suggested that the holotype of ''Mesadactylus'', Brigham Young University Museum of Paleontology, BYU 2024, a synsacrum, belonged to an anurognathid, though this affinity has been questioned by other authors. ''Mesadactylus'' is from the Late Jurassic Morrison Formation of the United States. Indeterminate anurognathid remains have also been reported from the Middle Jurassic Bakhar Svita of Mongolia and the Early Cretaceous of North Korea. Classification A family A ...
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Dendrorhynchoides
''Dendrorhynchoides'' was a genus of anurognathid pterosaur containing only the holotype species ''D. curvidentatus'' that is known from the Middle Jurassic Tiaojishan Formation of Qinglong, northern Hebei Province, China. The genus was first named '' Dendrorhynchus'' in 1998 by Ji Shu'an and Ji Qiang, but that name proved to be preoccupied by a parasitic protozoan named in 1920 by David Keilin. It was therefore renamed in 1999. The type species is ''Dendrorhynchoides curvidentatus''. The genus name is derived from Greek ''dendron'', "tree" and ''rhynkhos'', "snout" in reference to it being assumed a tree-dweller and presumed a close relative of ''Rhamphorhynchus''. The specific name means "curved-toothed" in Latin. A second species, ''D. mutoudengensis'', was described in 2012, and moved to a new genus ''Luopterus'' in 2020. Discovery and naming The genus is based on holotype GMV2128, a fossil originally discovered around 1995 and obtained by science from illegal fossil deale ...
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Jeholopterus
''Jeholopterus'' was a small anurognathid pterosaur from the Middle to Late Jurassic Daohugou Beds of the Tiaojishan Formation of Inner Mongolia, China, preserved with hair-like pycnofibres and skin remains. Naming The genus was named in 2002 by Wang Xiaolin, Zhou Zhonghe and Xu Xing. The type species, the only known, is ''Jeholopterus ninchengensis''. The genus name is derived from its place of discovery, Jehol in China, and a Latinized Greek ''pteron'', "wing". The specific name refers to Nincheng County. Description The type species is based on holotype IVPP V12705, a nearly complete specimen from the Daohugou beds of Ningcheng County in the Neimongol (Inner Mongolia) Autonomous Region of China.Wang, X., Zhou, Z., Zhang, F., and Xu, X. (2002). "A nearly completely articulated rhamphorhynchoid pterosaur with exceptionally well-preserved wing membranes and 'hairs' from Inner Mongolia, northeast China." ''Chinese Science Bulletin'' 47(3), 226 – 232. The specimen is cr ...
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Sinomacrops
''Sinomacrops'' is a genus of extinct anurognathid pterosaur from the Middle to Late Jurassic periods of what is now the Daohugou Beds of the Tiaojishan Formation in Mutoudeng, Qinglong County of the Hebei province. The remains of ''Sinomacrops'' date back to around 164 to 158 million years ago. The type and only known species is ''Sinomacrops bondei''. Etymology ''Sinomacrops'' derives from the Ancient Greek word roots ''Sino~'', referring to China, ''macro~'' (''makros''), meaning large, and ''ops'', meaning eyes/face. The name ''Sinomacrops'' is in reference to both the large eyes of and the broad faces that are typical of the family Anurognathidae, as well as to the Chinese origin of the animal. The specific name, ''bondei'', honors paleontologist Niels Bonde. Description ''Sinomacrops'' exhibits two autapomorphies (distinguishing traits) that distinguish it from other pterosaurs: one of them is having the first three maxillary alveoli (tooth sockets) closely spaced, and ...
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Anurognathus
''Anurognathus'' is a genus of small pterosaur that lived during the Late Jurassic period (Tithonian stage). ''Anurognathus'' was first named and described by Ludwig Döderlein in 1923.Döderlein, L. (1923). "''Anurognathus Ammoni'', ein neuer Flugsaurier". ''Sitzungsberichte der Mathematisch-Naturwissenschaftlichen Abteilung der Bayerischen Akademie der Wissenschaften zu München'', 1923, 306-307. The type species is ''Anurognathus ammoni''. The genus name ''Anurognathus'' is derived from the Greek ''αν''/ ''an-'' ("without"), ''оυρα''/ ''oura'' ("tail"), and ''γναθος''/ ''gnathos'' ("jaw") in reference to its unusually small tail relative to other "rhamphorhynchoid" (i.e. basal) pterosaurs."Anurognathus." In: Cranfield, Ingrid (ed.). ''The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures''. London: Salamander Books, Ltd. Pp. 292-295. The specific name ''ammoni'' honours the Bavarian geologist Ludwig von Ammon, from whose collection Döderlein had ...
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Anatoly Nicolaevich Ryabinin
Anatoliy Riabinin (11 June 1874-2 February 1942) was a Russian geologist and vertebrate paleontologist. In the 1910s, Riabinin led the first paleontological expeditions to the Amur (in 1914 and 1916–17). The hadrosaurid '' Amurosaurus riabinini'' is named for him. In 1925, he described the first dinosaur from China (though excavated in 1914). It initially bore the name ''Trachodon amurense'', though he renamed it '' Mandschurosaurus'' in 1930. He also conducted studies on fossilised turtles. In 2020, the ornithopod dinosaur genus known as ''Riabininohadros'' was named for him. In 1942, he perished during the Siege of Leningrad The siege of Leningrad (russian: links=no, translit=Blokada Leningrada, Блокада Ленинграда; german: links=no, Leningrader Blockade; ) was a prolonged military blockade undertaken by the Axis powers against the Soviet city of L .... Several of his works were posthumously published in the late 1940s (publications appeared in 1945 and 19 ...
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Karabastau Formation
The Karabastau Formation ( kk, Qarabastaý svıtasy) is a geological formation and lagerstätte in the Karatau Mountains of southern Kazakhstan whose strata date to the Middle to Late Jurassic. It is an important locality for insect fossils that has been studied since the early 20th century, alongside the rarer remains of vertebrates, including pterosaurs, salamanders, lizards and crocodiles.Barrett, P.M., Butler, R.J., Edwards, N.P., & Milner, A.R. Pterosaur distribution in time and space: an atlas. p61-107. in Flugsaurier: Pterosaur papers in honour of Peter Wellnhofer. 2008. Hone, D.W.E., and Buffetaut, E. (eds). Zitteliana B, 28. 264p/ref> Lithology and depositional environment The primary lithology consists of 1 mm thick varve laminations of claystone, with a dark part and a light dolomitic part, which probably correspond to a wet and dry season respectively, alongside rare, several cm thick sandstone interbeds. These were deposited within an ancient freshwater paleol ...
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Anatoly Riabinin
Anatoliy Riabinin (11 June 1874-2 February 1942) was a Russian geologist and vertebrate paleontologist. In the 1910s, Riabinin led the first paleontological expeditions to the Amur (in 1914 and 1916–17). The hadrosaurid '' Amurosaurus riabinini'' is named for him. In 1925, he described the first dinosaur from China (though excavated in 1914). It initially bore the name ''Trachodon amurense'', though he renamed it ''Mandschurosaurus'' in 1930. He also conducted studies on fossilised turtles. In 2020, the ornithopod dinosaur genus known as ''Riabininohadros'' was named for him. In 1942, he perished during the Siege of Leningrad. Several of his works were posthumously published in the late 1940s (publications appeared in 1945 and 1946). He thus described ''Batrachognathus ''Batrachognathus'' is an extinct genus of anurognathid pterosaur from the Late Jurassic ( Oxfordian to Kimmeridgian) Karabastau Formation of the central Asian republic of Kazakhstan. The genus was named in 1948 ...
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Late Jurassic
The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name "Malm" indicates rocks of Late Jurassic age. In the past, ''Malm'' was also used to indicate the unit of geological time, but this usage is now discouraged to make a clear distinction between lithostratigraphic and geochronologic/chronostratigraphic units. Subdivisions The Late Jurassic is divided into three ages, which correspond with the three (faunal) stages of Upper Jurassic rock: Paleogeography During the Late Jurassic Epoch, Pangaea broke up into two supercontinents, Laurasia to the north, and Gondwana to the south. The result of this break-up was the spawning of the Atlantic Ocean. However, at this time, the Atlantic Ocean was relatively narrow. Life forms of the epoch This epoch is well known for many famous types of dinosau ...
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Holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering. A holotype is not necessarily "typ ...
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Bavaria
Bavaria ( ; ), officially the Free State of Bavaria (german: Freistaat Bayern, link=no ), is a state in the south-east of Germany. With an area of , Bavaria is the largest German state by land area, comprising roughly a fifth of the total land area of Germany. With over 13 million inhabitants, it is second in population only to North Rhine-Westphalia, but due to its large size its population density is below the German average. Bavaria's main cities are Munich (its capital and largest city and also the third largest city in Germany), Nuremberg, and Augsburg. The history of Bavaria includes its earliest settlement by Iron Age Celtic tribes, followed by the conquests of the Roman Empire in the 1st century BC, when the territory was incorporated into the provinces of Raetia and Noricum. It became the Duchy of Bavaria (a stem duchy) in the 6th century AD following the collapse of the Western Roman Empire. It was later incorporated into the Holy Roman Empire, became an ind ...
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