Aysheaia
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Aysheaia
''Aysheaia'' is an extinct genus of soft-bodied lobopod, known from the middle Cambrian of North America, with an average body length of 1–6 cm. Anatomy ''Aysheaia'' has ten body segments, each of which has a pair of spiked, annulate legs. The animal is segmented, and looks somewhat like a bloated caterpillar with a few spines added on — including six finger-like projections around the mouth and two grasping legs on the "head". Each leg has a subterminal row of about six curved claws. No jaw apparatus is evident. A pair of legs marks the posterior end of the body, unlike in onychophorans where the anus projects posteriad; this may be an adaptation to the terrestrial habit. Ecology Based on its association with sponge remains, it is believed that ''Aysheaia'' was a sponge grazer and may have protected itself from predators by seeking refuge within sponge colonies. ''Aysheaia'' probably used its claws to cling to sponges. A terminal mouth is also seen in tardigrades th ...
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Aysheaia Pedunculata
''Aysheaia'' is an extinct genus of soft-bodied lobopod, known from the middle Cambrian of North America, with an average body length of 1–6 cm. Anatomy ''Aysheaia'' has ten body segments, each of which has a pair of spiked, annulate legs. The animal is segmented, and looks somewhat like a bloated caterpillar with a few spines added on — including six finger-like projections around the mouth and two grasping legs on the "head". Each leg has a subterminal row of about six curved claws. No jaw apparatus is evident. A pair of legs marks the posterior end of the body, unlike in onychophorans where the anus projects posteriad; this may be an adaptation to the terrestrial habit. Ecology Based on its association with sponge remains, it is believed that ''Aysheaia'' was a sponge grazer and may have protected itself from predators by seeking refuge within sponge colonies. ''Aysheaia'' probably used its claws to cling to sponges. A terminal mouth is also seen in tardigrades th ...
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Aysheaia2
''Aysheaia'' is an extinct genus of soft-bodied lobopod, known from the middle Cambrian of North America, with an average body length of 1–6 cm. Anatomy ''Aysheaia'' has ten body segments, each of which has a pair of spiked, annulate legs. The animal is segmented, and looks somewhat like a bloated caterpillar with a few spines added on — including six finger-like projections around the mouth and two grasping legs on the "head". Each leg has a subterminal row of about six curved claws. No jaw apparatus is evident. A pair of legs marks the posterior end of the body, unlike in onychophorans where the anus projects posteriad; this may be an adaptation to the terrestrial habit. Ecology Based on its association with sponge remains, it is believed that ''Aysheaia'' was a sponge grazer and may have protected itself from predators by seeking refuge within sponge colonies. ''Aysheaia'' probably used its claws to cling to sponges. A terminal mouth is also seen in tardigrades th ...
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Lobopodia
The lobopodians, members of the informal group Lobopodia (from the Greek language, Greek, meaning "blunt feet"), or the formally erected phylum Lobopoda Cavalier-Smith (1998), are panarthropods with stubby legs called lobopods, a term which may also be used as a common name of this group as well. While the definition of lobopodians may differ between literatures, it usually refers to a group of soft-bodied, worm-like fossil panarthropods such as ''Aysheaia'' and ''Hallucigenia''. The oldest near-complete fossil lobopodians date to the Lower Cambrian; some are also known from Ordovician, Silurian and Carboniferous Lagerstätten. Some bear toughened claws, plates or spines, which are commonly preserved as small carbonaceous fossil, carbonaceous or Small Shelly Fossils, mineralized microfossils in Cambrian strata. The grouping is considered to be paraphyletic, as the three living panarthropod groups (Arthropoda, Tardigrada and Onychophora) are thought to have evolved from lobopodian ...
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Stanleycaris
''Stanleycaris'' is an extinct, monotypic genus of hurdiid radiodont from the middle Cambrian (Miaolingian). The type species is ''Stanleycaris hirpex''. ''Stanleycaris'' was described from the Stephen Formation near the Stanley Glacier and Burgess Shale locality of Canada, as well as Wheeler Formation of United States. The genus was characterized by the rake-like frontal appendages with robust inner spines. ''Stanleycaris'' was originally described only from frontal appendages and oral cone. However, in 2022, 268 specimens of ''Stanleycaris'', many of which were complete, were studied, making ''Stanleycaris'' a well documented radiodont. ''Stanleycaris'' had three eyes, a bizarre configuration previously unknown among other radiodont genera; yet this head anatomy supports early differentiation among arthropod head and trunk segmentation. The original description of the taxon appeared in an online supplement to the article published by Jean-Bernard Caron, Robert R. Gaines, M. ...
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Burgess Shale Fossils
The fossils of the Burgess Shale, like the Burgess Shale itself, formed around 505 million years ago in the Mid Cambrian period. They were discovered in Canada in 1886, and Charles Doolittle Walcott collected over 65,000 specimens in a series of field trips up to the alpine site from 1909 to 1924. After a period of neglect from the 1930s to the early 1960s, new excavations and re-examinations of Walcott's collection continue to reveal new species, and statistical analysis suggests that additional discoveries will continue for the foreseeable future. Stephen Jay Gould's book '' Wonderful Life'' describes the history of discovery up to the early 1980s, although his analysis of the implications for evolution has been contested. The fossil beds are in a series of shale layers, averaging and totalling about in thickness. These layers were deposited against the face of a high undersea limestone cliff. All these features were later raised up above current sea level during the cr ...
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Tardigrade
Tardigrades (), known colloquially as water bears or moss piglets, are a phylum of eight-legged segmented micro-animals. They were first described by the German zoologist Johann August Ephraim Goeze in 1773, who called them Kleiner Wasserbär ("little water bear"). In 1777, the Italian biologist Lazzaro Spallanzani named them Tardigrada (), which means "slow steppers". They have been found in diverse regions of Earth's biospheremountaintops, the deep sea, tropical rainforests, and the Antarctic. Tardigrades are among the most resilient animals known, with individual species able to survive extreme conditions – such as exposure to extreme temperatures, extreme pressures (both high and low), air deprivation, radiation, dehydration, and starvation – that would quickly kill most other known forms of life. Tardigrades have survived exposure to outer space. There are about 1,300 known species in the phylum Tardigrada, a part of the superphylum Ecdysozoa consisting of animals th ...
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Radiodont
Radiodonta is an extinct Order (biology), order of stem-group arthropods that was successful worldwide during the Cambrian period. They may be referred to as radiodonts, radiodontans, radiodontids, anomalocarids, or anomalocaridids, although the last two originally refer to the family Anomalocarididae, which previously included all species of this order but is now restricted to only a few species. Radiodonts are distinguished by their distinctive frontal appendages, which are morphologically diverse and used for a variety of functions. Radiodonts included the earliest large predators known, but they also included sediment sifters and filter feeders. Some of the most famous species of radiodonts are the Cambrian taxa ''Anomalocaris, Anomalocaris canadensis'', ''Hurdia, Hurdia victoria'', ''Peytoia nathorsti'', ''Titanokorys gainesii, Titanokorys gainessii, Cambroraster, Cambroraster falcatus'' and ''Amplectobelua, Amplectobelua symbrachiata'', the Ordovician ''Aegirocassis, Aegiroca ...
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Wheeler Formation
The Wheeler Shale (named by Charles Walcott) is a Cambrian ( 507  Ma) fossil locality world-famous for prolific agnostid and ''Elrathia kingii'' trilobite remains (even though many areas are barren of fossils) and represents a Konzentrat-Lagerstätte. Varied soft bodied organisms are locally preserved, a fauna (including ''Naraoia'', ''Wiwaxia'' and ''Hallucigenia'') and preservation style (carbonaceous film) normally associated with the more famous Burgess Shale. As such, the Wheeler Shale also represents a Konservat-Lagerstätten. Together with the Marjum Formation and lower Weeks Formation, the Wheeler Shale forms of limestone and shale exposed in one of the thickest, most fossiliferous and best exposed sequences of Middle Cambrian rocks in North America. At the type locality of Wheeler Amphitheater, House Range, Millard County, western Utah, the Wheeler Shale consists of a heterogeneous succession of highly calcareous shale, shaley limestone, mudstone and thi ...
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Wheeler Shale
The Wheeler Shale (named by Charles Walcott) is a Cambrian ( 507  Ma) fossil locality world-famous for prolific agnostid and ''Elrathia kingii'' trilobite remains (even though many areas are barren of fossils) and represents a Konzentrat-Lagerstätte. Varied soft bodied organisms are locally preserved, a fauna (including ''Naraoia'', ''Wiwaxia'' and ''Hallucigenia'') and preservation style (carbonaceous film) normally associated with the more famous Burgess Shale. As such, the Wheeler Shale also represents a Konservat-Lagerstätten. Together with the Marjum Formation and lower Weeks Formation, the Wheeler Shale forms of limestone and shale exposed in one of the thickest, most fossiliferous and best exposed sequences of Middle Cambrian rocks in North America. At the type locality of Wheeler Amphitheater, House Range, Millard County, western Utah, the Wheeler Shale consists of a heterogeneous succession of highly calcareous shale, shaley limestone, mudstone and thi ...
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Phyllopod Bed
The Phyllopod bed, designated by USNM locality number 35k, is the most famous fossil-bearing member of the Burgess Shale fossil ''Lagerstätte''. It was quarried by Charles Walcott from 1911–1917 (and later named Walcott Quarry), and was the source of 95% of the fossils he collected during this time; tens of thousands of soft-bodied fossils representing over 150 genera have been recovered from the Phyllopod bed alone. Stratigraphy and location The phyllopod bed is a 2.31 m thick layer of the 7 m thick Greater Phyllopod Bed, found in the Walcott Quarry on Fossil Ridge, between Wapta Mountain and Mount Field, at an elevation of around , around north of the railway town of Field, British Columbia, in the Canadian Rocky Mountains. It is adjacent to Mount Burgess, where Walcott first discovered the Burgess Shale formation. Walcott divided the bed into twelve units based on the rock type and fossil content. Certain fossil beds provide reference levels and can b ...
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Xenusion
''Xenusion auerswaldae'' is an early lobopodian known from two specimens found in glacial erratics on the Baltic coast of Germany. They probably originated in the Kalmarsund Sandstone of Southern Sweden, which was deposited in the Lower Cambrian (Upper Tommotian–Lower Atdabanian; Stages 2→3). It is the oldest currently known lobopodian with soft body fossils. The specimens are not especially well preserved. The older specimen is 10 cm or so in length with a narrow, weakly segmented body. Assuming it was the posterior section, the specimen was estimated to be part of an animal about 20 cm in length. A depression runs up the bottom on all but the rearmost segments. There is a slightly bulbous termination, and each segment before that seems to have a single pair of tapering annulated legs similar to the modern onychophoran, but without specialized feet and claws. More than 10 body segments were present. There is presumably a spine on each body bump and faint trans ...
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Onychophora
Onychophora (from grc, ονυχής, , "claws"; and , , "to carry"), commonly known as velvet worms (due to their velvety texture and somewhat wormlike appearance) or more ambiguously as peripatus (after the first described genus, '' Peripatus''), is a phylum of elongate, soft-bodied, many-legged panarthropods. In appearance they have variously been compared to worms with legs, caterpillars, and slugs. They prey upon other invertebrates, which they catch by ejecting an adhesive slime. Approximately 200 species of velvet worms have been described, although the true number of species is likely greater. The two extant families of velvet worms are Peripatidae and Peripatopsidae. They show a peculiar distribution, with the peripatids being predominantly equatorial and tropical, while the peripatopsids are all found south of the equator. It is the only phylum within Animalia that is wholly endemic to terrestrial environments, at least among extant members. Velvet worms are generally c ...
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