Avicularium (bivalve)
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Avicularium (bivalve)
The avicularium (pl. avicularia) in Cheilostomata, cheilostome bryozoans is a modified, non-feeding zooid. The Operculum (bryozoa), operculum, which normally closes the orifice when the zooids tentacles are retracted, has been modified to become a mandible. Strong muscles operate it. The polypide is greatly reduced, and the individual receives nourishment from neighboring zooids. The shape of the avicularian zooid can be identical to the feeding autozooid, but is usually elongated in the direction of the mandible. Avicularia are categorised by their position relative to the autozooids. Vicarious avicularia effectively replace an autozooid in the colony structure and are usually a similar size and shape as the autozooids. Interzooidal avicularia are wedged between autozooids but do not replace an autozooid. Adventitious avicularia are placed somewhere on the external (frontal) wall of an autozooid and are usually much smaller. The 'birds-head' avicularium (e.g. ''Bugula'') is elevated ...
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Cheilostomata
Cheilostomatida, also called Cheilostomata, is an order of Bryozoa in the class Gymnolaemata. They are exclusively marine, colonial invertebrate animals. Cheilostome colonies are composed of calcium carbonate and grow on a variety of surfaces, including rocks, shells, seagrass and kelps. The colony shapes range from simple encrusting sheets to erect branching and even unattached forms. As in other bryozoan groups, each colony is composed of a few to thousands of individual polypides. Each individual has a U-shaped gut, and no respiratory, circulatory, or nerve system. Unique among bryozoans, cheilostome polypides are housed in a box-shaped zooids, which do not grow larger once the zooid is mature. The opening through which the polypide protrudes is protected by a calcareous or chitinous lidlike structure, an operculum. Cheilostomes possess avicularia, which have modified the operculum into a range of mandibles (possibly for defense) or hair-like setae (possibly for cleaning). ...
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Bryozoa
Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic invertebrate animals, nearly all living in sedentary colonies. Typically about long, they have a special feeding structure called a lophophore, a "crown" of tentacles used for filter feeding. Most marine bryozoans live in tropical waters, but a few are found in oceanic trenches and polar waters. The bryozoans are classified as the marine bryozoans (Stenolaemata), freshwater bryozoans (Phylactolaemata), and mostly-marine bryozoans (Gymnolaemata), a few members of which prefer brackish water. 5,869living species are known. At least two genera are solitary (''Aethozooides'' and ''Monobryozoon''); the rest are colonial. The terms Polyzoa and Bryozoa were introduced in 1830 and 1831, respectively. Soon after it was named, another group of animals was discovered whose filtering mechanism looked similar, so it was included in Bryozoa until 1869, when the two groups were no ...
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Zooid
A zooid or zoöid is a single animal that is part of a colonial animal. This lifestyle has been adopted by animals from separate unrelated taxa. Zooids are multicellular; their structure is similar to that of other solitary animals. The zooids can either be directly connected by tissue (e.g. corals, Catenulida, Siphonophorae, Pyrosome or Ectoprocta) or share a common exoskeleton (e.g. Bryozoa or Pterobranchia). The colonial organism as a whole is called a ''zoon'' , plural ''zoa'' (from Ancient Greek meaning animal; plural , ). Zooids can exhibit polymorphism. For instance, extant bryozoans may have zooids adapted for different functions, such as feeding, anchoring the colony to the substratum and for brooding embryos. However, fossil bryozoans are only known by the colony structures that the zooids formed during life. There are correlations between the size of some zooids and temperature. Variations in zooid size within colonies of fossils can be used as an indicator of ...
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Operculum (bryozoa)
In the bryozoan order Cheilostomatida, the operculum is a calcareous or chitinous lid-like structure that protects the opening through which the polypide protrudes. Many species have modified the operculum in specialized zooids (avicularia) to form a range of mandibles (probably for defense) or hair-like setae (probably for cleaning, or in some unattached species, such as '' Selenaria'', for locomotion). The cyclostome family Eleidae also convergently evolved an opercular structure during the Early Cretaceous to Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''pal ....Taylor, P.D. 1994. Systematics of the melicerititid cyclostome bryozoans; introduction and the genera ''Elea'', ''Semielea'' and ''Repromultelea''. Bulletin of the Natural history Museum, Geology Series 50:1 ...
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Polypide
The polypide in bryozoans encompasses most of the organs and tissues of each individual zooid. This includes the tentacles, tentacle sheath, U-shaped digestive tract, musculature and nerve cells. It is housed in the zooidal exoskeleton, which in cyclostomes Cyclostome is a biological term (from the Greek for "round mouth") used in a few different senses: * for the taxon Cyclostomi, which comprises the extant jawless fishes: the hagfish (Myxini) and the lampreys (Petromyzontidae). This was thought fo ... is tubular and in cheilostomes is box-shaped. See also Bryozoan Anatomy References Bryozoology Protostome anatomy {{bryozoan-stub ...
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Bugula
''Bugula'' is a genus of common colonial arborescent bryozoa, often mistaken for seaweed. It commonly grows upright in bushy colonies of up to 15 cm in height.Bock, P. (2015). Bugula Oken, 1815. In: Bock, P.; Gordon, D. (2015) World List of Bryozoa. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=110839 on 2015-09-02 Distribution The native distribution of ''Bugula neritina'' is presumed to be tropical and subtropical waters; however, it has become widespread globally due to attachment to the hulls of vessels. It is considered an invasive species in some countries. ''Bugula neritina'' ''Bugula neritina'' attracted interest as a source of cytotoxic chemicals, bryostatins, under clinical investigation as anti-cancer agents. It was first collected and provided to JL Hartwell's anticancer drug discovery group at the National Cancer Institute (NCI) by Jack Rudloe. In 2001 pharmaceutical company GPC Biotech licensed ...
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Vibraculum
Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic invertebrate animals, nearly all living in sedentary colonies. Typically about long, they have a special feeding structure called a lophophore, a "crown" of tentacles used for filter feeding. Most marine bryozoans live in tropical waters, but a few are found in oceanic trenches and polar waters. The bryozoans are classified as the marine bryozoans (Stenolaemata), freshwater bryozoans (Phylactolaemata), and mostly-marine bryozoans (Gymnolaemata), a few members of which prefer brackish water. 5,869living species are known. At least two genera are solitary (''Aethozooides'' and ''Monobryozoon''); the rest are colonial. The terms Polyzoa and Bryozoa were introduced in 1830 and 1831, respectively. Soon after it was named, another group of animals was discovered whose filtering mechanism looked similar, so it was included in Bryozoa until 1869, when the two groups were not ...
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Cyclostomatida
Cyclostomatida, or cyclostomata (also known as cyclostomes), are an ancient order of stenolaemate bryozoans which first appeared in the Lower Ordovician. It consists of 7+ suborders, 59+ families, 373+ genera, and 666+ species. The cyclostome bryozoans were dominant in the Mesozoic; since that era, they have decreased. Currently, cyclostomes seldom constitute more than 20% of the species recorded in regional bryozoan faunas. Taxonomy Traditionally, cyclostomes have been divided into two groups according to the skeletal organization. In free-walled (or double-walled) cyclostomes, the exterior frontal walls of the zooids are uncalcified; autozooids have either a polygonal aperture bounded by vertical interior walls, or a subcircular aperture in species with kenozooids filling the spaces between the autozooids. By contrast, fixed-walled (or single-walled) cyclostomes have much of the exterior frontal wall calcified; autozooids normally have a subcircular aperture located at or ...
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Convergent Evolution
Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last common ancestor of those groups. The cladistic term for the same phenomenon is homoplasy. The recurrent evolution of flight is a classic example, as flying insects, birds, pterosaurs, and bats have independently evolved the useful capacity of flight. Functionally similar features that have arisen through convergent evolution are ''analogous'', whereas '' homologous'' structures or traits have a common origin but can have dissimilar functions. Bird, bat, and pterosaur wings are analogous structures, but their forelimbs are homologous, sharing an ancestral state despite serving different functions. The opposite of convergence is divergent evolution, where related species evolve different traits. Convergent evolution is similar to parallel evo ...
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Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of the entire Phanerozoic. The name is derived from the Latin ''creta'', "chalk", which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation ''Kreide''. The Cretaceous was a period with a relatively warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now- extinct marine reptiles, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was ice free, and forests extended to the poles. During this time, new groups of mammals and birds appeared. During the Early Cretaceous, flowering plants appeared and began to rapidly diversify, becoming the dominant group of plants across the Earth b ...
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