Atlantic Horseshoe Crab
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Atlantic Horseshoe Crab
The Atlantic horseshoe crab (''Limulus polyphemus''), also known as the American horseshoe crab, is a species of marine and brackish chelicerate arthropod. Despite their name, horseshoe crabs are more closely related to spiders, ticks, and scorpions than to crabs. It is found in the Gulf of Mexico and along the Atlantic coast of North America. The main area of annual migration is Delaware Bay along the South Jersey Delaware Bayshore. Their eggs were eaten by Native Americans, but today Atlantic horseshoe crabs are caught for use as fishing bait, in biomedicine (especially for ''Limulus'' amebocyte lysate) and science. They play a major role in the local ecosystems, with their eggs providing an important food source for shorebirds, and the juveniles and adults being eaten by sea turtles. The other three extant (living) species in the family Limulidae are also called horseshoe crabs, but they are restricted to Asia. Names and classification This group of animals is also known ...
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Carl Linnaeus
Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the modern system of naming organisms. He is known as the "father of modern taxonomy". Many of his writings were in Latin; his name is rendered in Latin as and, after his 1761 ennoblement, as . Linnaeus was born in Råshult, the countryside of Småland, in southern Sweden. He received most of his higher education at Uppsala University and began giving lectures in botany there in 1730. He lived abroad between 1735 and 1738, where he studied and also published the first edition of his ' in the Netherlands. He then returned to Sweden where he became professor of medicine and botany at Uppsala. In the 1740s, he was sent on several journeys through Sweden to find and classify plants and animals. In the 1750s and 1760s, he continued to collect an ...
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Limulus Amebocyte Lysate
Limulus amebocyte lysate (LAL) is an aqueous extract of blood cells (amoebocytes) from the Atlantic horseshoe crab ''Limulus polyphemus''. LAL reacts with bacterial endotoxin lipopolysaccharide (LPS), which is a membrane component of gram-negative bacteria. This reaction is the basis of the LAL test, which is widely used for the detection and quantification of bacterial endotoxins. In Asia, a similar Tachypleus amebocyte lysate (TAL) test based on the local horseshoe crabs ''Tachypleus gigas'' or ''Tachypleus tridentatus'' is occasionally used instead. The recombinant factor C (rFC) assay is a replacement of LAL/TAL based on a similar reaction. Background The American medical researcher Fred Bang reported in 1956 that gram-negative bacteria, even if killed, will cause the blood of the horseshoe crab to turn into a semi-solid mass. It was later recognized that the animal's blood cells, mobile cells called amoebocytes, contain granules with a clotting factor known as coaguloge ...
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Yucatán Peninsula
The Yucatán Peninsula (, also , ; es, Península de Yucatán ) is a large peninsula in southeastern Mexico and adjacent portions of Belize and Guatemala. The peninsula extends towards the northeast, separating the Gulf of Mexico to the north and west of the peninsula from the Caribbean Sea to the east. The Yucatán Channel, between the northeastern corner of the peninsula and Cuba, connects the two bodies of water. The peninsula is approximately in area. It has low relief, and is almost entirely composed of porous limestone. The peninsula lies east of the Isthmus of Tehuantepec, the narrowest point in Mexico separating the Atlantic Ocean, including the Gulf of Mexico and Caribbean Sea, from the Pacific Ocean. Some consider the isthmus to be the geographic boundary between Central America and the rest of North America, placing the peninsula in Central America. Politically all of Mexico, including the Yucatán, is generally considered part of North America, while Guatemala an ...
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Microsatellite DNA
A microsatellite is a tract of repetitive DNA in which certain DNA motifs (ranging in length from one to six or more base pairs) are repeated, typically 5–50 times. Microsatellites occur at thousands of locations within an organism's genome. They have a higher mutation rate than other areas of DNA leading to high genetic diversity. Microsatellites are often referred to as short tandem repeats (STRs) by forensic geneticists and in genetic genealogy, or as simple sequence repeats (SSRs) by plant geneticists. Microsatellites and their longer cousins, the minisatellites, together are classified as VNTR (variable number of tandem repeats) DNA. The name "satellite" DNA refers to the early observation that centrifugation of genomic DNA in a test tube separates a prominent layer of bulk DNA from accompanying "satellite" layers of repetitive DNA. They are widely used for DNA profiling in cancer diagnosis, in kinship analysis (especially paternity testing) and in forensic identificat ...
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Polyphemus
Polyphemus (; grc-gre, Πολύφημος, Polyphēmos, ; la, Polyphēmus ) is the one-eyed giant son of Poseidon and Thoosa in Greek mythology, one of the Cyclopes described in Homer's ''Odyssey''. His name means "abounding in songs and legends". Polyphemus first appeared as a savage man-eating giant in the ninth book of the ''Odyssey''. The satyr play of Euripides is dependent on this episode apart from one detail; Polyphemus is made a pederast in the play. Later Classical writers presented him in their poems as heterosexual and linked his name with the nymph Galatea. Often he was portrayed as unsuccessful in these, and as unaware of his disproportionate size and musical failings. In the work of even later authors, however, he is presented as both a successful lover and skilled musician. From the Renaissance on, art and literature reflect all of these interpretations of the giant. Odysseus and Polyphemus Ancient sources In Homer's epic, Odysseus lands on the island of the ...
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Decapoda
The Decapoda or decapods (literally "ten-footed") are an order of crustaceans within the class Malacostraca, including many familiar groups, such as crabs, lobsters, crayfish, shrimp and prawns. Most decapods are scavengers. The order is estimated to contain nearly 15,000 species in around 2,700 genera, with around 3,300 fossil species. Nearly half of these species are crabs, with the shrimp (about 3,000 species) and Anomura including hermit crabs, porcelain crabs, squat lobsters (about 2500 species) making up the bulk of the remainder. The earliest fossil decapod is the Devonian ''Palaeopalaemon''. Anatomy Decapods can have as many as 38 appendages, arranged in one pair per body segment. As the name Decapoda (from the Greek , ', "ten", and , '' -pod'', "foot") implies, ten of these appendages are considered legs. They are the pereiopods, found on the last five thoracic segments. In many decapods, one pair of these "legs" has enlarged pincers, called chelae, with the legs be ...
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Telmessus Cheiragonus
Telmessos or Telmessus ( Hittite: 𒆪𒉿𒆷𒉺𒀸𒊭 ''Kuwalapašša'',  Lycian: 𐊗𐊁𐊍𐊁𐊂𐊁𐊛𐊆 ''Telebehi'', grc, Τελμησσός), also Telmissus ( grc, Τελμισσός), later Anastasiopolis ( grc, Αναστασιούπολις), then Makri or Macre ( el, Μάκρη), was the largest city in Lycia, near the Carian border, and is sometimes confused with Telmessos in Caria. It was called Telebehi in the Lycian language. The well-protected harbor of Telmessos is separated from the Gulf of Telmessos by an island. The name of the modern town on the site is Fethiye. History The city was known as Kuwalapašša by the Hittites and Telebehi by the Lycians. According to the annals of Hattusili III, the city was a part of Lycia (which was known as Lukka at the time) and conquered by the Hittites. Another Hittite document mentions the cities of Kuwalapašša and Dalawa sent aid to Hittites during the war against Iyalanda. Telmessos was a flourishin ...
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Malacostracan
Malacostraca (from New Latin; ) is the largest of the six classes of crustaceans, containing about 40,000 living species, divided among 16 orders. Its members, the malacostracans, display a great diversity of body forms and include crabs, lobsters, crayfish, shrimp, krill, prawns, woodlice, amphipods, mantis shrimp, tongue-eating lice and many other less familiar animals. They are abundant in all marine environments and have colonised freshwater and terrestrial habitats. They are segmented animals, united by a common body plan comprising 20 body segments (rarely 21), and divided into a head, thorax, and abdomen. Etymology The name Malacostraca was coined by a French zoologist Pierre André Latreille in 1802. He was curator of the arthropod collection at the National Museum of Natural History in Paris. The name comes from the Greek roots (', meaning "soft") and (', meaning "shell"). The name is misleading, since the shell is soft only immediately after moulting, and is usuall ...
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Atlantic Horseshoe Crab (Limulus Polyphemus) (5)
The Atlantic horseshoe crab (''Limulus polyphemus''), also known as the American horseshoe crab, is a species of horseshoe crab, a kind of marine and brackish chelicerate arthropod. It is found in the Gulf of Mexico and along the Atlantic coast of North America. The main area of annual migration is Delaware Bay along the South Jersey Delaware Bayshore. Their eggs were eaten by Native Americans, but today Atlantic horseshoe crabs are caught for use as fishing bait, in biomedicine (especially for ''Limulus'' amebocyte lysate) and science. They play a major role in the local ecosystems, with their eggs providing an important food source for shorebirds, and the juveniles and adults being eaten by sea turtles. The other three extant (living) species in the family Limulidae are also called horseshoe crabs, but they are restricted to Asia. Names and classification This group of animals is also known as horsefoot, or saucepan. Some people call the horseshoe crab a "helmet crab", but ...
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Horseshoe Crab With Shells
A horseshoe is a fabricated product designed to protect a horse hoof from wear. Shoes are attached on the palmar surface (ground side) of the hooves, usually nailed through the insensitive hoof wall that is anatomically akin to the human toenail, although much larger and thicker. However, there are also cases where shoes are glued. Horseshoes are available in a wide variety of materials and styles, developed for different types of horse and for the work they do. The most common materials are steel and aluminium, but specialized shoes may include use of rubber, plastic, magnesium, titanium, or copper.Price, Steven D. (ed.) ''The Whole Horse Catalog: Revised and Updated'' New York:Fireside 1998 , pp. 84–87. Steel tends to be preferred in sports in which a strong, long-wearing shoe is needed, such as polo, eventing, show jumping, and western riding events. Aluminium shoes are lighter, making them common in horse racing where a lighter shoe is desired, and often facilitate certa ...
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Limulidae
Horseshoe crabs are marine and brackish water arthropods of the family Limulidae and the only living members of the order Xiphosura. Despite their name, they are not true crabs or crustaceans: they are chelicerates, most closely related to arachnids, such as spiders and scorpions. Horseshoe crabs live primarily in and around shallow coastal waters on soft, sandy or muddy bottoms. They are generally found in the intertidal zone at spring high tides. They are eaten in some parts of Asia, and used as fishing bait, in fertilizer and in science (especially ''Limulus'' amebocyte lysate). In recent years, population declines have occurred as a consequence of coastal habitat destruction and overharvesting. Tetrodotoxin may be present in one horseshoe crab species, '' Carcinoscorpius rotundicauda''. Fossil records for horseshoe crabs extend back as far as 480 million years ago, with extant forms being living fossils. A 2019 molecular analysis places them as the sister group of Ricinulei ...
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Extant Taxon
Neontology is a part of biology that, in contrast to paleontology, deals with living (or, more generally, '' recent'') organisms. It is the study of extant taxa (singular: extant taxon): taxa (such as species, genera and families) with members still alive, as opposed to (all) being extinct. For example: * The moose (''Alces alces'') is an extant species, and the dodo (''Raphus cucullatus'') is an extinct species. * In the group of molluscs known as the cephalopods, there were approximately 600 extant species and 7,500 extinct species. A taxon can be classified as extinct if it is broadly agreed or certified that no members of the group are still alive. Conversely, an extinct taxon can be reclassified as extant if there are new discoveries of living species ("Lazarus species"), or if previously-known extant species are reclassified as members of the taxon. Most biologists, zoologists, and botanists are in practice neontologists, and the term neontologist is used large ...
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