Athyriaceae
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Athyriaceae
The Athyriaceae (ladyferns and allies) are a family of terrestrial ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the family is placed in the suborder Aspleniineae, and includes two genera. Alternatively, it may be treated as the subfamily Athyrioideae of a very broadly defined family Aspleniaceae. The family has with a cosmopolitan distribution. Description Species of the Athyriaceae are terrestrial or lithophytic, less commonly aquatic. They grow from various kinds of rhizome: short or long, creeping or erect, branched or not. The distribution and evolution of characters in the family is complex, and the genera have few constant features by which they can be identified. The sporangia have stalks two or three cells wide in the middle, and contain brown monolete spores. Taxonomy Earlier classifications The family was first created by Arthur H.G. Alston in 1956. It has had a varied history. In 2014, Christenhusz ...
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Athyriaceae
The Athyriaceae (ladyferns and allies) are a family of terrestrial ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the family is placed in the suborder Aspleniineae, and includes two genera. Alternatively, it may be treated as the subfamily Athyrioideae of a very broadly defined family Aspleniaceae. The family has with a cosmopolitan distribution. Description Species of the Athyriaceae are terrestrial or lithophytic, less commonly aquatic. They grow from various kinds of rhizome: short or long, creeping or erect, branched or not. The distribution and evolution of characters in the family is complex, and the genera have few constant features by which they can be identified. The sporangia have stalks two or three cells wide in the middle, and contain brown monolete spores. Taxonomy Earlier classifications The family was first created by Arthur H.G. Alston in 1956. It has had a varied history. In 2014, Christenhusz ...
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Polypodiales
The order Polypodiales encompasses the major lineages of polypod ferns, which comprise more than 80% of today's fern species. They are found in many parts of the world including tropical, semitropical and temperate areas. Description Polypodiales are unique in bearing sporangia with a vertical annulus interrupted by the stalk and stomium. These sporangial characters were used by Johann Jakob Bernhardi to define a group of ferns he called the "Cathetogyratae"; the Pteridophyte Phylogeny Group has suggested reviving this name as the informal term cathetogyrates, to replace the ambiguously circumscribed term "polypods" when referring to the Polypodiales. The sporangia are born on stalks 1–3 cells thick and are often long-stalked. (In contrast, the Hymenophyllales have a stalk composed of four rows of cells.) The sporangia do not reach maturity simultaneously. Many groups in the order lack indusia, but when present, they are attached either along the edge of the indusium or in its ...
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Diplazium
''Diplazium'' is a genus of ferns that specifically includes the approximately 400 known species of twinsorus ferns. The Greek root is ''diplazein'' meaning ''double'': the indusia in this genus lie on both sides of the vein. These ferns were earlier considered part of either the Athyriaceae, Dryopteridaceae, Aspleniaceae, or Polypodiaceae families or recognized as belonging to their own taxonomic family. The Pteridophyte Phylogeny Group classification of 2016 (PPG I) places the genus in the Athyriaceae. The taxonomy of the genus is difficult and poorly known, and by 2009 has never been the subject of a complete monographic study. Their distribution is pantropical, with a few species extending into temperate areas. The rhizome of the genus ''Diplazium'' varies from creeping to erect, and is scaly. Its fronds are deciduous or evergreen, are trophopodicThe trophopod is food storage organ described from a number of North American ferns. It consists of the enlarged and modified ...
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Athyrium
''Athyrium'' (lady-fern) is a genus of about 180 species of terrestrial ferns, with a cosmopolitan distribution. It is placed in the family Athyriaceae, in the order Polypodiales. Its genus name is from Greek '' a-'' ('without') and Latinized Greek ''thyreos'' ('shield'), describing its inconspicuous indusium (sorus' covering). The common name "lady fern" refers in particular to the common lady fern, ''Athyrium filix-femina''.Entry "lady fern", ''New Oxford American Dictionary 3rd edition'' (2010) by Oxford University Press, Inc. ''Athyrium'' species are used as food plants by the larvae of some Lepidoptera species including the small angle shades and ''Sthenopis auratus ''Sthenopis pretiosus'', the gold-spotted ghost moth, is a species of moth of the family Hepialidae. It was first described by Gottlieb August Wilhelm Herrich-Schäffer in 1856. It can be found in found Brazil, Venezuela and in the eastern United ...''. Species There are about 180, including: References ...
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Aspleniineae
Aspleniineae is a suborder of ferns in the order Polypodiales. It is equivalent to the clade eupolypods II in earlier systems; it is also treated as a single very broadly defined family Aspleniaceae. The suborder generally corresponds with the order Blechnales as described by J. L. Reveal in 1993. Aspleniineae includes some important ferns, including ''Onoclea sensibilis'', the sensitive fern, which grows as a virtual weed throughout much of its temperate North American range, and ferns of the genus ''Thelypteris'', a genus that has shown remarkable speciation. It also includes one of the more common horticultural ferns, ''Matteuccia struthiopteris'', the ostrich fern. Taxonomy In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the group is treated as the suborder Aspleniinae, and divided into 11 families. Alternatively, it may be treated as a single, very broadly circumscribed family Aspleniaceae ''sensu lato ''Sensu'' is a Latin word meaning "in the sense o ...
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Deparia
''Deparia'' (or the false spleenworts) is a genus of ferns. The Pteridophyte Phylogeny Group classification of 2016 (PPG I) places the genus in the family Athyriaceae, although other sources include it within an expanded Aspleniaceae or Woodsiaceae. Species , the ''Checklist of Ferns and Lycophytes of the World'' accepted the following species (with a note that there are "probably still too many described species in China"). *'' Deparia abbreviata'' (W.M.Chu) Z.R.He *''Deparia acrostichoides ''Deparia acrostichoides'', commonly called silvery glade fern or silvery spleenwort, is a perennial species of fern. Its range include much of the eastern United States and Canada, from Ontario to Nova Scotia, and Georgia to Louisiana and easter ...'' (Sw.) M.Kato *'' Deparia allantodioides'' (Bedd.) M.Kato *'' Deparia auriculata'' (W.M.Chu & Z.R.Wang) Z.R.Wang *'' Deparia biserialis'' (Baker) M.Kato *'' Deparia bonincola'' (Nakai) M.Kato *'' Deparia boryana'' (Willd.) M.Kato *'' Deparia ...
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Fern
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except the lycopods, and differ from mosses and other bryophytes by being vascular, i.e., having specialized tissues that conduct water and nutrients and in having life cycles in which the branched sporophyte is the dominant phase. Ferns have complex leaves called megaphylls, that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns. They produce coiled fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae) and eusporangiate ferns, the latter group including horsetails, whisk ferns, marattioid ferns, and ophioglossoid ferns. Ferns first ...
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Pteridophyte Phylogeny Group
The Pteridophyte Phylogeny Group, or PPG, is an informal international group of systematic botanists who collaborate to establish a consensus on the classification of pteridophytes (lycophytes and ferns) that reflects knowledge about plant relationships discovered through phylogenetic studies. In 2016, the group published a classification for extant pteridophytes, termed "PPG I". The paper had 94 authors (26 principal and 68 additional). PPG I A first classification, PPG I, was produced in 2016, covering only extant (living) pteridophytes. The classification was rank-based, using the ranks of class, subclass, order, suborder, family, subfamily and genus. Phylogeny The classification was based on a consensus phylogeny, shown below to the level of order. The very large order Polypodiales was divided into two suborders, as well as families not placed in a suborder: Classification to subfamily level To the level of subfamily, the PPG I classification is as follows. *Class Lycopodi ...
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Aspleniaceae
The Aspleniaceae (spleenworts) are a family of ferns, included in the order Polypodiales. The composition and classification of the family have been subject to considerable changes. In particular, there is a narrow circumscription, Aspleniaceae s.s. (adopted here), in which the family contains only two genera, and a very broad one, Aspleniaceae s.l., in which the family includes 10 other families kept separate in the narrow circumscription, with the Aspleniaceae s.s. being reduced to the subfamily Asplenioideae. The family has a worldwide distribution, with many species in both temperate and tropical areas. Elongated unpaired sori are an important characteristic of most members of the family. Description Members of the family grow from rhizomes, that are either creeping or somewhat erect, and are usually but not always unbranched, and have scales that usually have a lattice-like (clathrate) structure. In some species, for example ''Asplenium nidus'', the rhizomes form a kind o ...
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Woodsiaceae
''Woodsia'' is a genus of ferns in the order Polypodiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), it is the only genus in the family Woodsiaceae, placed in the suborder Aspleniineae. The family can also be treated as the subfamily Woodsioideae of a very broadly defined family Aspleniaceae ''Sensu, sensu lato''. Species of ''Woodsia'' are commonly known as cliff ferns. Taxonomy Woodsiaceae formerly included the members of the families Athyriaceae and Diplaziopsidaceae, but analysis has consistently shown that they should be treated as separate families. The following cladogram for the suborder Aspleniineae (as eupolypods II), based on Lehtonen (2011), and Rothfels & al. (2012), shows a likely phylogenetics, phylogenetic relationship between the Woodsiaceae and the other families of the Aspleniineae. Species There are about 40–50 species of the genus ''Woodsia''. , ''Plants of the World Online'' accepted the following species: *''Woodsia alpi ...
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PPG I
The Pteridophyte Phylogeny Group, or PPG, is an informal international group of systematic botanists who collaborate to establish a consensus on the classification of pteridophytes (lycophytes and ferns) that reflects knowledge about plant relationships discovered through phylogenetic studies. In 2016, the group published a classification for extant pteridophytes, termed "PPG I". The paper had 94 authors (26 principal and 68 additional). PPG I A first classification, PPG I, was produced in 2016, covering only extant (living) pteridophytes. The classification was rank-based, using the ranks of class, subclass, order, suborder, family, subfamily and genus. Phylogeny The classification was based on a consensus phylogeny, shown below to the level of order. The very large order Polypodiales was divided into two suborders, as well as families not placed in a suborder: Classification to subfamily level To the level of subfamily, the PPG I classification is as follows. *Class Lycopo ...
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Circumscription (taxonomy)
In biological taxonomy, circumscription is the content of a taxon, that is, the delimitation of which subordinate taxa are parts of that taxon. If we determine that species X, Y, and Z belong in Genus A, and species T, U, V, and W belong in Genus B, those are our circumscriptions of those two genera. Another systematist might determine that T, U, V, W, X, Y, and Z all belong in genus A. Agreement on circumscriptions is not governed by the Codes of Zoological or Botanical Nomenclature, and must be reached by scientific consensus. A goal of biological taxonomy is to achieve a stable circumscription for every taxon. This goal conflicts, at times, with the goal of achieving a natural classification that reflects the evolutionary history of divergence of groups of organisms. Balancing these two goals is a work in progress, and the circumscriptions of many taxa that had been regarded as stable for decades are in upheaval in the light of rapid developments in molecular phylogenetics ...
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