Astroviida
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Astroviida
Lituitida is an order of orthoceratoid cephalopods. They correspond to the family Lituitidae of the Treatise (Furnish & Glenister, 1964), reranked as an order and combined with other orthoceratoids. They are considered to be more closely related to the Orthocerida than to the Ascocerida or Pseudorthocerida which are also included. Lituitids are characterized by smooth to annulate shells in which the juvenile portion near the apex is coiled or cyrtoconic. The adult portion is straight or slightly sigmoidal and may be expanded. Growth lines show a deep hyponomic sinus and lateral ocular sinuses. The siphuncle is subcentral with layered connecting rings. Cameral deposits are prevalent and concentrated more ventrally. The apex is small and spherical and lacks a cicatrix. The Lituitida, which were originally included in the Tarphycerida as the Lituitidae (Furnish and Glenister) have their beginnings in the Early Ordovician with forms like ''Ancistroceras''. Their diversity increased af ...
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Orthoceratoidea
Orthoceratoidea is a major subclass of nautiloid cephalopods. Members of this subclass usually have orthoconic (straight) to slightly cyrtoconic (curved) shells, and central to subcentral siphuncles which may bear internal deposits. Orthoceratoids are also characterized by dorsomyarian muscle scars (a small number of large scars concentrated at the top of the body chamber), extensive cameral deposits, and calciosiphonate connecting rings with a porous and calcitic inner layer. Currently, Orthoceratoidea comprises the orders Riocerida, Dissidocerida, Actinocerida, Pseudorthocerida, Lituitida and Orthocerida. Orthocerida is a noteworthy paraphyletic order which is ancestral to the major cephalopod groups such as the extinct ammonoids and living coleoids (cephalopods without external shells, including squids, octopus, cuttlefish, etc.). Taxonomy As a superorder, Orthoceratiodea was one of six superorders within the Nautiloidea, the others being the Plectronoceratoidea (= Ellesm ...
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Cephalopod
A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by ''Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been ident ...
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Orthocerida
Orthocerida is an order of extinct Orthoceratoid cephalopods also known as the Michelinocerida that lived from the Early Ordovician () possibly to the Late Triassic (). A fossil found in the Caucasus suggests they may even have survived until the Early Cretaceous (). They were most common however from the Ordovician to the Devonian. Shell form The shell is usually long, and may be straight ("orthoconic") or gently curved. In life, these animals may have been similar to the modern squid, except for the long shell. The internal structure of the shell consists of concavo-convex chambers linked by a centrally-placed tube called a siphuncle. There is a tendency for the chambers to develop cameral deposits, which were used as ballast to balance the long gas-filled shell. Depending on the family, the siphuncle has orthochoanitic (short and straight) or cyrtochoanitic (outwardly curved) septal necks, which protrude from the septa. The shell surface may be (depending on the specie ...
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Ascocerida
The Ascocerida are comparatively small, bizarre Orthoceratoids known only from Ordovician and Silurian sediments in Europe and North America, uniquely characterized by a deciduous conch consisting of a longiconic juvenile portion and an inflated breviconic adult portion that separate sometime in maturity.W.M . Furnish and Brian F Glenister; Nautiloidea-Ascocerida, K261-K277; Treatise on Invertebrate Paleontology , part K, Nautiloidea; Geological Society of America and the University of Kansas Press, 1964 Morphology The juvenile portion of an ascocerid consists typically of a narrow, circular cyrtocone that underwent periodic truncation. Cumulative length, including broken off segments, may have reached about in the largest individuals. The siphuncle is located half way between the shell axis and the venter, is thin walled and tubular with short, orthochoanitic septal necks and segments that are only slightly inflated. The juvenile portion is known as the deciduous conch, as it ...
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Pseudorthocerida
Pseudorthocerida is an order of generally straight longiconic orthoceratoids with a subcentral to marginal cyrtochoanitic siphuncle composed of variably expanded segments which may contain internal deposits that may develop into a continuous parietal lining. (Sweet 1964). Cameral deposits are common and concentrated ventrally. Apices typically have a slight to moderate exogastric curvature The Pseudorthocerida are included in a broad in-group of generally orthoconic cephalopods known as the Orthoceratoidea (Kroger 2008) along with the Ascocerida, Dissidocerida, Lituitida, and Orthocerida. The Pseudorthocerida were among the last living orthoconic nautiloids. One family, the Trematoceratidae, survived into the Triassic Period. Taxonomy Current understanding By current understanding the Pseudorthocerida contains the following families. : Pseudorthoceratidae : Cayutoceratidae : Pseudactinoceratidae : Spyroceratidae : Carbactinoceratidae : Trematoceratidae The Pseudorth ...
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Connecting Ring
The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and ''Spirula''. In the case of the cuttlefish, the siphuncle is indistinct and connects all the small chambers of that animal's highly modified shell; in the other cephalopods it is thread-like and passes through small openings in the septa (walls) dividing the camerae (chambers). Some older studies have used the term siphon for the siphuncle, though this naming convention is uncommon in modern studies to prevent confusion with a mollusc organ of the same name. Function The siphuncle is used primarily in emptying water from new chambers as the shell grows. To perform this task, the cephalopod increases the saltiness of the blood in the siphuncle, and the water moves from the more dilute chamber into the blood through osmosis. At the sam ...
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Ancistroceras
''Ancistroceras'' is one of the two ancestral lituitids from the late Early Ordovician (Arenigian). The other being '' Holmiceras''. The shell is weakly annulate, starts off with 1.5 to 2 contiguous or slightly separated whorls followed by a rapidly expanding orthocone with an apical angle of about 30 deg. Growth lines are sinuous, show a pronounced hyponomic sinus in the coiled juvenile portion at the apex but almost none at the mature aperture of the orthcone. ''Holmiceras'' is similar except that its whorls are more loosely coiled and it is known only from the Arenigian whereas ''Ancistroceras'' ranges through most of the Middle Ordovician. ''Ancistroceras'' may have given rise to '' Rhynchorthoceras'' by a loss of the juvenile coiled portion, replacing it with a short semi-tubular cyrtoconic section. ''Ancistroceras'' may also have given rise to ''Lituites ''Lituites'' is an extinct nautiloid genus from the Middle Ordovician and type for the Lituitidae (a tarphycerid fa ...
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Canadian Epoch
The Canadian is the Lower or Early Ordovician in North America. The term is common in the older literature and has been well understood for more than a century. However it has no official recognition by the International Commission on Stratigraphy (ICS) and has been superseded by the more recently defined Ibexian series of western Utah. Background Dana introduced the Canadian as the name for a system separated from the rest of the Ordovician (Weller 1980), then known as the Lower Silurian, and referred to the rest of the Lower Silurian as the Trenton System. At that time the Ordovician had not yet been recognized. Later Ulrich redefined the Canadian as roughly equivalent to the Beekmantown strata of the Lower Ordovician. Flower (1957 p. 17) felt that recognition of the Canadian as a separate system would greatly solve problems in Early Paleozoic stratigraphy. As such, faunas in limestones of Canadian age are uniformly widespread and set off sharply from black shale graptolite ...
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Lituites
''Lituites'' is an extinct nautiloid genus from the Middle Ordovician and type for the Lituitidae (a tarphycerid family) that in some more recent taxonomies has been classified with the orthocerids and listed under the order Lituitida. Fossils have been found in New York, Argentina, Norway, Sweden, Estonia, and China. ''Lituites'' produced a shell with a planospirally coiled juvenile portion at the apex, reflective of its tarphycerid ancestry, followed by a long, moderately expanding, generally straight, orthoconic adult section with a subdorsal siphuncle connecting the chambers. The adult body chamber may equal or exceed the length of the chambered part of the orthoconic section. The mature aperture has a pair of pronounced ventrolateral lappets and a similar but shorter pair of dorsolateral lappets. ''Lituites'' gives its name to the term "lituiticone" which refers to a shell that is coiled in the early growth stage and later becomes uncoiled. References *Flower & ...
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Cephalopod Orders
A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by ''Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been identified. Tw ...
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