Asteroxylales
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Asteroxylales
Drepanophycales is an order of extinct lycopsid plants of Late Silurian to Late Devonian age (around ), found in North America, China, Russia, Europe, and Australia. Sometimes known as the Asteroxylales or Baragwanathiales. Description Extinct terrestrial vascular plants of the Silurian to Devonian periods. Stem of the order of several mm to several cm in diameter and several cm to several metres long, erect or arched, dichotomizing occasionally, furnished with true roots at the base. Vascular bundle an exarch actinostele, tracheids of primitive annular or helical type (so-called G-type). Stem clothed in either microphylls (leaves with a single vascular thread or 'vein'), or with leaf-like enations (unvascularized projections) with a vascular trace into the base of each enation. Homosporous, with sporangia borne singly and dehiscing by a single slit. List of families The following families have, at various times, been segregated within the Drepanophycales. However, Kenrick ...
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Drepanophycales
Drepanophycales is an order of extinct lycopsid plants of Late Silurian to Late Devonian age (around ), found in North America, China, Russia, Europe, and Australia. Sometimes known as the Asteroxylales or Baragwanathiales. Description Extinct terrestrial vascular plants of the Silurian to Devonian periods. Stem of the order of several mm to several cm in diameter and several cm to several metres long, erect or arched, dichotomizing occasionally, furnished with true roots at the base. Vascular bundle an exarch actinostele, tracheids of primitive annular or helical type (so-called G-type). Stem clothed in either microphylls (leaves with a single vascular thread or 'vein'), or with leaf-like enations (unvascularized projections) with a vascular trace into the base of each enation. Homosporous, with sporangia borne singly and dehiscing by a single slit. List of families The following families have, at various times, been segregated within the Drepanophycales. However, Kenrick and ...
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Lycopodiopsida
Lycopodiopsida is a class of vascular plants known as lycopods, lycophytes or other terms including the component lyco-. Members of the class are also called clubmosses, firmosses, spikemosses and quillworts. They have dichotomously branching stems bearing simple leaves called microphylls and reproduce by means of spores borne in sporangia on the sides of the stems at the bases of the leaves. Although living species are small, during the Carboniferous, extinct tree-like forms formed huge forests that dominated the landscape and contributed to coal deposits. The nomenclature and classification of plants with microphylls varies substantially among authors. A consensus classification for extant (living) species was produced in 2016 by the Pteridophyte Phylogeny Group (PPG I), which places them all in the class Lycopodiopsida, which includes the classes Isoetopsida and Selaginellopsida used in other systems. (See Table 2.) Alternative classification systems have used ranks fro ...
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Asteroxylon
''Asteroxylon'' ("star-shaped xylem") is an extinct genus of vascular plants of the Division Lycopodiophyta known from anatomically preserved specimens described from the famous Early Devonian Rhynie chert and Windyfield chert in Aberdeenshire, Scotland. ''Asteroxylon'' is considered a basal member of the Lycopsida. Description ''Asteroxylon'' is a genus of terrestrial vascular plant which flourished in the Early Devonian period. This plant consisted of aerial, isotomously and anisotomously branching stems that reached 12 mm in diameter and 40 cm in length. The possibly procumbent aerial stems arose from a leaf-less rhizome which bore smaller-diameter, positively geotropic root-like branches. The rhizomes, which represent an independent origin of roots, reached a depth of up to 20 cm below the surface. A 407 million-year-old fossil from the Rhynie chert shows the roots formed through a modified version of a mechanism called “dichotomous branching”, where one ...
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Sporangium
A sporangium (; from Late Latin, ) is an enclosure in which spores are formed. It can be composed of a single cell or can be multicellular. Virtually all plants, fungi, and many other lineages form sporangia at some point in their life cycle. Sporangia can produce spores by mitosis, but in nearly all land plants and many fungi, sporangia are the site of meiosis and produce genetically distinct haploid spores. Fungi In some phyla of fungi, the sporangium plays a role in asexual reproduction, and may play an indirect role in sexual reproduction. The sporangium forms on the sporangiophore and contains haploid nuclei and cytoplasm. Spores are formed in the sporangiophore by encasing each haploid nucleus and cytoplasm in a tough outer membrane. During asexual reproduction, these spores are dispersed via wind and germinate into haploid hyphae. Although sexual reproduction in fungi varies between phyla, for some fungi the sporangium plays an indirect role in sexual reprod ...
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Silurian First Appearances
The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods (myriapods, arachnids and hexapods) became fully terrestrialized. A significant evolutionary milestone during th ...
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Silurian Plants
The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods (myriapods, arachnids and hexapods) became fully terrestrialized. A significant evolutionary milestone during th ...
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Devonian Plants
The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, where rocks from this period were first studied. The first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents. By the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. The arthropod groups of myriapods, arachnids and hexapods also became well-established early in this period, after starting their expansion to land at least from the Ordovician period. Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The placoderms began dominating al ...
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Baragwanathia
''Baragwanathia'' is a genus of extinct lycopsid plants of Late Silurian to Early Devonian age (), fossils of which have been found in Australia, Canada, China and Czechia. The name derives from William Baragwanath who discovered the first specimens of the type species, ''Baragwanathia longifolia'', at Thomson River (Victoria, Australia). Description ''Baragwanathia'' differed from such taxa as '' Asteroxylon'' by the presence of vascular tissue in its leaves—''Asteroxylon'' had enations without vascular tissue. The sporangia were borne in the axils of the leaves, which were spirally arranged. By comparison, the closely related genus '' Drepanophycus'' of the same period (see Drepanophycaceae for more details) bore its sporangia on the upper surface of specialized leaves known as sporophylls. ''Baragwanathia'' varied in size, with stems up to a few cm in diameter and up to a few metres in length. They were erect or arched, dichotomized (forked) occasionally, and had advent ...
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Drepanophycus
''Drepanophycus'' is a genus of extinct plants of the division Lycopodiophyta of Early to Late Devonian age (around ), found in Eastern Canada and Northeast US, China, Russia, Egypt and various parts of Northern Europe and Britain. Description Extinct terrestrial vascular plants of the Devonian period. Stem of the order of several mm to several cm in diameter and several cm to a metre long, erect or arched, dichotomizing occasionally, furnished with true roots at the base. Vascular bundle actinostele, tracheids of primitive annular or helical type (so-called G-type). Leaves are unbranched thorn-shaped (i.e. with a wide base, tapering to a blunt point) microphylls several mm long with a single prominent vascular thread, arranged spirally to randomly on the stem. Sporangia borne singly on the upper leaf surface. ''Drepanophycus'' has similarities to the genus ''Halleophyton''. It differs from a closely related genus of the same period, Baragwanathia, in the position of the sporang ...
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Microphyll
In plant anatomy and evolution a microphyll (or lycophyll) is a type of plant leaf with one single, unbranched leaf vein. Plants with microphyll leaves occur early in the fossil record, and few such plants exist today. In the classical concept of a microphyll, the leaf vein emerges from the protostele without leaving a leaf gap. Leaf gaps are small areas above the node of some leaves where there is no vascular tissue, as it has all been diverted to the leaf. Megaphylls, in contrast, have multiple veins within the leaf and leaf gaps above them in the stem. Leaf vasculature The clubmosses and horsetails have microphylls, as in all extant species there is only a single vascular trace in each leaf. These leaves are narrow because the width of the blade is limited by the distance water can efficiently diffuse cell-to-cell from the central vascular strand to the margin of the leaf. Despite their name, microphylls are not always small: those of ''Isoëtes'' can reach 25 centimetres ...
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Asteroxylaceae
''Asteroxylon'' ("star-shaped xylem") is an extinct genus of vascular plants of the Division Lycopodiophyta known from anatomically preserved specimens described from the famous Early Devonian Rhynie chert and Windyfield chert in Aberdeenshire, Scotland. ''Asteroxylon'' is considered a basal member of the Lycopsida. Description ''Asteroxylon'' is a genus of terrestrial vascular plant which flourished in the Early Devonian period. This plant consisted of aerial, isotomously and anisotomously branching stems that reached 12 mm in diameter and 40 cm in length. The possibly procumbent aerial stems arose from a leaf-less rhizome which bore smaller-diameter, positively geotropic root-like branches. The rhizomes, which represent an independent origin of roots, reached a depth of up to 20 cm below the surface. A 407 million-year-old fossil from the Rhynie chert shows the roots formed through a modified version of a mechanism called “dichotomous branching”, where ...
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Tracheid
A tracheid is a long and tapered lignified cell in the xylem of vascular plants. It is a type of conductive cell called a tracheary element. Angiosperms use another type of tracheary element, called vessel elements, to transport water through the xylem. The main functions of tracheid cells are to transport water and inorganic salts, and to provide structural support for trees. There are often pits on the cell walls of tracheids, which allows for water flow between cells. Tracheids are dead at functional maturity and do not have a protoplast. The wood (softwood) of gymnosperms such as pines and other conifers is mainly composed of tracheids. Tracheids are also the main conductive cells in the primary xylem of ferns. The tracheid was first named after the German botanist Carl Gustav Sanio in 1863, from the German ''Tracheide''. Evolution Tracheids were the main conductive cells found in early vascular plants. In the first 140-150 million years of vascular plant evolution, tr ...
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