Articulata (Brachiopoda)
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Articulata (Brachiopoda)
Rhynchonelliformea is a major subphylum and clade of brachiopods. It is equivalent to the former class Articulata, which was used previously in brachiopod taxonomy. Articulate brachiopods have many anatomical differences relative to "inarticulate" brachiopods of the subphyla Linguliformea and Craniformea. Articulates have hard calcium carbonate shells with tongue-and-groove hinge articulations (hence the name) and separate sets of simple opening and closing muscles. The name, Rhynchoelliformea, which replaces Articulata, which is also a class of crinoids, comes from the family Rhynchonellidae, which however is no more representative of articulate brachiopods than any other group such as spirifers or strophomenids. It just happens to be the name chosen, based on an included taxon. The main difference between the Rhynchonelliformea described in the Treatise Part H, revised 2000/2007, and the Articulata of the Treatise part H, 1965, lies in the groups included, their taxonomic po ...
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Lower Cambrian
The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Cambrian biolo ...
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Phylogeny
A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological species or other entities based upon similarities and differences in their physical or genetic characteristics. All life on Earth is part of a single phylogenetic tree, indicating common ancestry. In a ''rooted'' phylogenetic tree, each node with descendants represents the inferred most recent common ancestor of those descendants, and the edge lengths in some trees may be interpreted as time estimates. Each node is called a taxonomic unit. Internal nodes are generally called hypothetical taxonomic units, as they cannot be directly observed. Trees are useful in fields of biology such as bioinformatics, systematics, and phylogenetics. ''Unrooted'' trees illustrate only the relatedness of the leaf nodes and do not require the ancestral root to be ...
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Athyridida
Athyridida is an order of Paleozoic brachiopods included in the Rhynchonellata, which makes up part of the articulate brachiopods. The Athyridida are the Rostrospracea of R.C Moore, 1952,Moore, Lalcker and Fischer, 1952, Invertebrate Fossils, McGraw-Hill considered at that time to be a suborder of the Spiriferida. As with the Spiriferida, the Athyridida have outwardly directed spiral brachidia that support the lophophores on either side, but instead have non-plicate shells with rounded outlines and prominent beaks but almost no inner areas on the pedicle valve. Athyridids began early in the Silurian, reached their greatest diversity in the following Devonian, and from then declined steadily until almost becoming extinct at the end of the Permian The order rejuvenated somewhat during the Triassic, only to decline again until becoming extinct in the Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (strat ...
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Paleozoic
The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ''zōḗ'' (), "life", meaning "ancient life" ). It is the longest of the Phanerozoic eras, lasting from , and is subdivided into six geologic periods (from oldest to youngest): # Cambrian # Ordovician # Silurian # Devonian # Carboniferous # Permian The Paleozoic comes after the Neoproterozoic Era of the Proterozoic Eon and is followed by the Mesozoic Era. The Paleozoic was a time of dramatic geological, climatic, and evolutionary change. The Cambrian witnessed the most rapid and widespread diversification of life in Earth's history, known as the Cambrian explosion, in which most modern phyla first appeared. Arthropods, molluscs, fish, amphibians, reptiles, and synapsids all evolved during the Paleozoic. Life began in the ocean ...
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Treatise On Invertebrate Paleontology
The ''Treatise on Invertebrate Paleontology'' (or ''TIP'') published by the Geological Society of America and the University of Kansas Press, is a definitive multi-authored work of some 50 volumes, written by more than 300 paleontologists, and covering every phylum, class, order, family, and genus of fossil and extant (still living) invertebrate animals. The prehistoric invertebrates are described as to their taxonomy, morphology, paleoecology, stratigraphic and paleogeographic range. However, taxa with no fossil record whatsoever have just a very brief listing. Publication of the decades-long ''Treatise on Invertebrate Paleontology'' is a work-in-progress; and therefore it is not yet complete: For example, there is no volume yet published regarding the post-Paleozoic era caenogastropods (a molluscan group including the whelk and Common periwinkle, periwinkle). Furthermore, every so often, previously published volumes of the ''Treatise'' are revised. Evolution of the proje ...
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Strophomenida
Strophomenida is an extinct order of articulate brachiopods which lived from the lower Ordovician period to the mid Carboniferous period. Strophomenida is part of the extinct class Strophomenata, and was the largest known order of brachiopods, encompassing over 400 genera. Some of the largest and heaviest known brachiopod species belong to this class. Strophomenids were among the most diverse and abundant brachiopods during the Ordovician, but their diversity was strongly impacted at the Late Ordovician mass extinction. Survivors rediversified into new morphologies in the Silurian, only to be impacted once again at the Late Devonian mass extinction. However, they still survived till the mid Carboniferous. Adult strophomenids lack an opening for the pedicle (stalk), so in life, they either lay free or cemented the ventral valve (lower shell) onto a firm substrate at the umbo (hinge). In juveniles, a tiny hole for the pedicle was present on the ventral valve near the umbo, but ...
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Spiriferida
Spiriferida is an order of extinct articulate brachiopod fossils which are known for their long hinge-line, which is often the widest part of the shell. In some genera (e.g. ''Mucrospirifer'') it is greatly elongated, giving them a wing-like appearance. They often have a deep fold down the center of the shell. The feature that gives the spiriferids their name ("spiral-bearers") is the internal support for the lophophore; this ''brachidium'', which is often preserved in fossils, is a thin ribbon of calcite that is typically coiled tightly within the shell. Spiriferids first appear in the Late Ordovician with the appearance of ''Eospirifer radiatus''. They increased in diversity throughout the Silurian and underwent a dramatic evolutionary radiation during the Devonian period, reaching peak development in variety and numbers. Spiriferida survived the great Permian extinction, finally becoming extinct during the Early to Middle Jurassic. Fossils of this order are often preserved a ...
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Crinoidea
Crinoids are marine animals that make up the class Crinoidea. Crinoids that are attached to the sea bottom by a stalk in their adult form are commonly called sea lilies, while the unstalked forms are called feather stars or comatulids, which are members of the largest crinoid order, Comatulida. Crinoids are echinoderms in the phylum Echinodermata, which also includes the starfish, brittle stars, sea urchins and sea cucumbers. They live in both shallow water and in depths as great as . Adult crinoids are characterised by having the mouth located on the upper surface. This is surrounded by feeding arms, and is linked to a U-shaped gut, with the anus being located on the oral disc near the mouth. Although the basic echinoderm pattern of fivefold symmetry can be recognised, in most crinoids the five arms are subdivided into ten or more. These have feathery pinnules and are spread wide to gather planktonic particles from the water. At some stage in their lives, most crinoids have ...
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