Anhangueria
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Anhangueria
Anhangueria (or anhanguerians) is a group of pterosaurs belonging to the clade Pteranodontoidea. Fossil remains of this group date back from the Early to Late Cretaceous periods (Valanginian to Turonian stages), around 140 to 92.5 million years ago. Anhangueria was named by paleontologists Taissa Rodrigues and Alexander Kellner in a review of ''Ornithocheirus'' species in 2013, they defined the clade as a branch-based taxon consisting of all pteranodontoids more closely related to ''Anhanguera blittersdorffi'' than to ''Istiodactylus latidens'' and ''Cimoliopterus cuvieri''. Classification Anhangueria originally only contained the genera '' Brasileodactylus'', ''Camposipterus'', ''Cearadactylus'', ''Ludodactylus'' as well as the family Anhangueridae, however, recent analyses had recovered the family Hamipteridae within this clade as well. In 2014, paleontologist Brian Andres and colleagues assigned more groups and genera within this clade, this included ''Guidraco'', the subfa ...
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Cimoliopterus Cuvieri
''Cimoliopterus'' is a genus of pterosaur that lived during the Late Cretaceous in what is now England and the United States. The first known specimen, consisting of the front part of a snout including part of a crest, was discovered in the Grey Chalk Subgroup of Kent, England, and described as the new species '' Pterodactylus cuvieri'' in 1851. The specific name ''cuvieri'' honours the palaeontologist George Cuvier, whereas the genus ''Pterodactylus'' was then used for many pterosaur species that are not thought to be closely related today. It was one of the first pterosaurs to be depicted as models in Crystal Palace Park in the 1850s. The species was subsequently assigned to various other genera, including '' Ornithocheirus'' and ''Anhanguera''. In 2013, the species was moved to a new genus, as ''Cimoliopterus cuvieri''; the generic name ''Cimoliopterus'' is derived from the Greek words for "chalk" and "wing". Other specimens and species have also been assigned to or synonym ...
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Ornithocheiridae
Ornithocheiridae (or ornithocheirids, meaning "bird hands") is a group of pterosaurs within the suborder Pterodactyloidea. These pterosaurs were among the last to possess teeth. Members that belong to this group lived from the Early to Late Cretaceous periods (Valanginian to Turonian stages), around 140 to 90 million years ago. Ornithocheirids are generally infamous for having an enormously controversial and very confusing taxonomy. Although agreements that these animals were related, and therefore similar to istiodactylids and pteranodontians, there is still no virtual consensus over the exact content and interrelationships of this group. Ornithocheirids were the most successful pterosaurs during their reign, they were also the largest pterosaurs before the appearance of the azhdarchids such as ''Quetzalcoatlus''. Ornithocheirids were excellent fish hunters, they used various flight techniques to catch their prey, and they are also capable of flying great distances without fl ...
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Lanceodontia
Ornithocheiromorpha (from Ancient Greek, meaning "bird hand form") is a group of pterosaurs within the suborder Pterodactyloidea. Fossil remains of this group date back from the Early to Late Cretaceous periods (Valanginian to Turonian stages), around 140 to 92.5 million years ago. Ornithocheiromorphs were discovered worldwide except Antarctica, though most genera were recovered in Europe, Asia and South America. They were the most diverse and successful pterosaurs during the Early Cretaceous, but throughout the Late Cretaceous they were replaced by better adapted and more advanced pterosaur species such the pteranodontids and azhdarchoids. The Ornithocheiromorpha was defined in 2014 by Andres and colleagues, and they made Ornithocheiromorpha the most inclusive clade containing ''Ornithocheirus'', but not '' Pteranodon''. Ornithocheiromorphs are considered to be some of the largest animals to have ever flown. Members of this group are also regarded to have some of the lar ...
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Iberodactylus
''Iberodactylus'' is a genus of pterodactyloid pterosaurs belonging to the clade Anhangueria, that during the Early Cretaceous lived in the area of present Spain. The type species is ''Iberodactylus andreui''. History of discovery and naming In the late 1980s, amateur paleontologist Javier Andreu discovered a pterosaur skull at the Los Quiñones site, west of Obón in Aragón. At the time it represented the most complete discovery of pterosaur fossil material in Spain apart from the remains of '' Prejanopterus''. In 2014, the find was reported in the scientific literature by José Antonio Ulloa-Rivas and identified as a member of the Ornithocheiroidea.Ulloa-Rivas, J.A. & Canudo, J.I. 2014. "New cranial remains of Ornithocheiroidea (Pterosauria) from the Barremian (Lower Cretaceous) of the Iberian Peninsula (Nuevos restos craneales de Ornithocheiroidea (Pterosauria) del Barremiense (Cretácico Inferior) de la Península Ibérica)". In: (Rodríguez, G.J.A. et alii, eds) ''New In ...
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Ornithocheirae
Ornithocheirae is an extinct clade of pteranodontoid pterosaurs from the Early Cretaceous to the Late Cretaceous (Valanginian to Turonian stages) of Asia, Europe, North America and South America. It was named by Harry Seeley in 1870 as a family that contains ''Ornithocheirus'' and its relatives. The name was emended to Ornithocheiridae, to match the requirements of the ICZN Code that a family-ranked clade should end with an -idae suffix. Brian Andres (2010) in his review of pterosaur phylogeny, defined the name Ornithocheirae phylogenetically, as a node-based taxon consisting of the last common ancestor of ''Anhanguera'' and ''Ornithocheirus'' and all its descendants. Thus Ornithocheirae is defined to include two families, the Anhangueridae and the Ornithocheiridae, following the opinion of Alexander KellnerKellner, A. W. A., (2003): Pterosaur phylogeny and comments on the evolutionary history of the group. pp. 105-137. — ''in'' Buffetaut, E. & Mazin, J.-M., (eds.): ''Evolu ...
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Cearadactylus
''Cearadactylus'' is a genus of large anhanguerid pterodactyloid pterosaur from the Romualdo Formation of Brazil, South America. Fossil remains of ''Cearadactylus'' dated back to the Albian stage of the Early Cretaceous period, about 112 million years ago. The only known species is ''C. atrox'', described and named in 1985 by Giuseppe Leonardi and Guido Borgomanero. The name refers to the Brazilian state Ceará, and combines this with Greek ''daktylos'', "finger", a reference to the wing finger of pterosaurs. The Latin ''atrox'' means "frightful", a reference to the fearsome dentition of the species. Discovery The holotype of ''Cearadactylus'' is MN 7019-V (formerly CB-PV-F-O93), which was discovered in the Romualdo Formation of the Santana Group in the Araripe plateau of northeastern Brazil. It consists of a single skull with a length of .Leonardi, G. & Borgomanero, G. (1985). "''Cearadactylus atrox'' nov. gen., nov. sp.: novo Pterosauria (Pterodactyloidea) da Chapada do Ararip ...
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Boreopterinae
Boreopteridae (meaning "northern wings") is a group of pterodactyloid pterosaurs from the Aptian-age Lower Cretaceous Yixian Formation of Liaoning, China. Classification In 2006, Lü Junchang and colleagues named the clade Boreopteridae for the clade containing the common ancestor of '' Boreopterus'' and '' Feilongus'' and all its descendants, which the authors reclassified as close relatives of the ornithocheirids. (''Feilongus'' had originally been considered a gallodactylid). Many possible boreopterids were subsequently described, one possible example being '' Aetodactylus'', which has been claimed to be similar to '' Boreopterus''. Originally considered close relatives of the ornithocheirids, many of these supposed boreopterids have been found to belong to other groups of the pterodactyloid lineage. In 2012, a phylogenetic analysis by Lü ''et al.'' divided the Boreopteridae into two subfamilies: Boreopterinae, comprising ''Boreopterus'' and ''Zhenyuanopterus'', and Mog ...
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Ludodactylus
''Ludodactylus'' (meaning "play finger") is a genus of anhanguerid pterodactyloid pterosaur from the Early Cretaceous period (Aptian stage) of what is now the Crato Formation of the Araripe Basin in Ceará, Brazil. The type and only species is ''L. sibbicki''. The generic name ''Ludodactylus'' refers to the fact that the animal had the combination of teeth and a ''Pteranodon''-like head crest, similar to many toy pterosaurs, and no such creature was known to exist until the discovery of ''Ludodactylus''. However, ''Ludodactylus'' is not the only pterosaur known to possess these features, its very close relative ''Caulkicephalus'' is another example. Etymology The genus was named by Eberhard Frey ''et al.'' in 2003 and contains one known species, ''Ludodactylus sibbicki''. The name is derived from Latin ''ludus'', "play" and Greek ''daktylos'', "finger". ''Ludus'' refers to the fact, long lamented by paleontologists, that many toy pterosaurs combined teeth with a ''Pteranodon''-l ...
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Hamipteridae
Hamipteridae (or hamipterids) is a small family of anhanguerian pterosaurs known from the Early Cretaceous of China and Spain. Classification The cladogram below follows the topology recovered by Pêgas ''et al.'' (2019). In the analysis, they assigned Hamipteridae as the sister taxon of the family Anhangueridae, both within the larger clade Anhangueria Anhangueria (or anhanguerians) is a group of pterosaurs belonging to the clade Pteranodontoidea. Fossil remains of this group date back from the Early to Late Cretaceous periods (Valanginian to Turonian stages), around 140 to 92.5 million years ....Rodrigo V. Pêgas, Borja Holgado & Maria Eduarda C. Leal (2019) On ''Targaryendraco wiedenrothi'' gen. nov. (Pterodactyloidea, Pteranodontoidea, Lanceodontia) and recognition of a new cosmopolitan lineage of Cretaceous toothed pterodactyloids, Historical Biology, References Pteranodontoids Prehistoric reptile families {{pterosaur-stub ...
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Camposipterus
''Camposipterus'' is a genus of pterodactyloid pterosaur from the Early Cretaceous of England. Fossil remains of ''Camposipterus'' dated back to the Early Cretaceous, about 112 million years ago. Discovery and naming In 1869, Harry Govier Seeley, based on a fossil found at Haslingfield, Cambridgeshire, named ''Ptenodactylus nasutus'',Seeley, H.G., 1869, ''Index to the fossil remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata arranged in the Woodwardian Museum of the University of Cambridge''. Deighton, Bell and Co., Cambridge, xxiii + 143 pp at the same time disclaiming the name which makes it invalid by modern standards. In 1870, Seeley had realized that the generic name ''Ptenodactylus'' had been preoccupied, so he renamed the species into ''Ornithocheirus nasutus''.Seeley, H.G., 1870, ''The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwar ...
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Guidraco
''Guidraco'' (Chin. ''gui'' (鬼) "malicious ghost" + Lat. ''draco'' "dragon") is an extinct genus of toothed pterodactyloid pterosaur known from the Early Cretaceous of Liaoning Province, northeast China. According to many recent studies, ''Guidraco'' is a member of the group Anhanguerinae, a subfamily belonging to the larger group Anhangueridae. Discovery ''Guidraco'' is known only from the holotype IVPP V17083, an articulated partial skeleton consisting of a nearly complete skull, lower jaws and a series of four, second to fifth, cervical vertebrae. It was collected at Sihedang near Lingyuan City in the Liaoning Province from the Jiufotang Formation, dating to the Aptian stage of the Early Cretaceous, about 120 million years ago. Etymology ''Guidraco'' was first described and named by Wang Xiaolin, Alexander W.A. Kellner, Jiang Shunxing and Cheng Xin in 2012 and the type species is ''Guidraco venator''. The generic name is derived from Chinese ''gui'' (鬼), "m ...
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Hamipterid
Hamipteridae (or hamipterids) is a small Family (biology), family of anhanguerian pterosaurs known from the Early Cretaceous of China and Spain. Classification The cladogram below follows the topology recovered by Pêgas ''et al.'' (2019). In the analysis, they assigned Hamipteridae as the sister taxon of the family Anhangueridae, both within the larger clade Anhangueria.Rodrigo V. Pêgas, Borja Holgado & Maria Eduarda C. Leal (2019) On ''Targaryendraco wiedenrothi'' gen. nov. (Pterodactyloidea, Pteranodontoidea, Lanceodontia) and recognition of a new cosmopolitan lineage of Cretaceous toothed pterodactyloids, Historical Biology, References

Pteranodontoids Prehistoric reptile families {{pterosaur-stub ...
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