Ancylotherium
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Ancylotherium
''Ancylotherium'' (from Greek, meaning "hooked beast") is an extinct genus of the family Chalicotheriidae, subfamily Schizotheriinae, endemic to Europe, Asia, and Africa during the Late Miocene-Early Pleistocene (9.0—1.8 mya), existing for approximately . Taxonomy ''Ancylotherium'' was named by Gaudry (1863) and was assigned to Chalicotheriidae by Carroll (1988); and to Schizotheriinae by Geraads et al. (2007). Description With some individuals reaching high at the shoulder and a weight of , ''Ancylotherium'' was relatively large, and was built rather like a goat. Individuals varied considerably in size, and the genus may have been sexually dimorphic, like many other chalicotheres. While it had the typical long forelimbs and short hind limbs of a chalicothere, like other schizotheriines, it did not walk on its knuckles. It was similar to the North American genus ''Moropus''. In at least some individuals of ''A. pentelicum'' the frontal bone of the skull is inflated to form a ...
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Chalicotheriidae
Chalicotheres (from Greek '' chalix'', "gravel" and '' therion'', "beast") are an extinct clade of herbivorous, odd-toed ungulate (perissodactyl) mammals that lived in North America, Eurasia, and Africa from the Middle Eocene until the Early Pleistocene, existing from 48.6 to 1.806 mya. They are one of the five major radiations of perissodactyls, with three groups living ( horses, plus the extinct paleotheres; rhinoceroses; tapirs), and two extinct (brontotheres and chalicotheres). Description Unlike modern perissodactyls, chalicotheres had clawed feet. They had longer forelimbs and shorter hind limbs, lower incisors that cropped food against a toothless pad in the upper jaw, low-crowned molar teeth, and were browsers on trees and shrubs throughout their history. They evolved in two different directions, which became separate subfamilies, the Schizotheriinae and the Chalicotheriinae. Schizotherine chalicotheres such as ''Moropus'' lived in a variety of forest, woodland, a ...
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Tylocephalonyx
''Tylocephalonyx'' (Greek: "knob" (tylos), "head" (kephalos), "claw/hoof" (onyx)) is an extinct chalicothere from the Miocene of North America. Description ''Tylocephalonyx'' specimens are notable for a dome-shaped skull, a feature found in some other schizotheriine chalicotheres but most developed in this genus. Adequate fossil material is lacking to tell whether both sexes had domed heads, but sexual dimorphism was common in the group. ''Tylocephalonyx'' may have used its "dome" in the same way as the pachycephalosaurs, though there is no clear evidence to link either pachycephalosaurs nor ''Tylocephalonyx'' to using their domes to crash together in high-impact head-to-head contests, as in modern bighorn sheep. Such contests require special cranial adaptations to protect the brain and cervical spine, not shown in chalicotheres. The dome of ''Tylocephalonyx'' may have been used for visual display or in butting or head-to-body battering contests. ''Tylocephalonyx'' is a perrisod ...
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Schizotheriinae
Schizotheriines are one of the two subfamilies of the extinct family Chalicotheriidae, a group of herbivorous odd-toed ungulate (perissodactyl) mammals that lived from the Eocene to the Pleistoscene. The other clade is the Chalicotheriinae. Both clades had claws rather than hooves on their front feet, an adaptation understood as related to feeding. Schizotheriines also had claws on their hind feet. The fossils of both groups are found in environments that had trees and shrubs. While chalicotheriines developed very derived body forms, schizotheriines remained basically similar in shape to other perissodactyls such as horses and tapirs. Like most forest-dwelling ungulates, they had long necks and forelimbs longer than their hindlimbs. Schizotheriines had longer, higher-crowned cheek teeth than chalicotheriines, which indicates they typically fed on tougher vegetation. The sediments where their fossils are found show they also lived in a wider range of environments, from moist forest ...
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Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the ...
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Fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the ...
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Rhinoceros
A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct species of the superfamily Rhinocerotoidea.) Two of the extant species are native to Africa, and three to South and Southeast Asia. Rhinoceroses are some of the largest remaining megafauna: all weigh at least one tonne in adulthood. They have a herbivorous diet, small brains (400–600 g) for mammals of their size, one or two horns, and a thick (1.5–5 cm), protective skin formed from layers of collagen positioned in a lattice structure. They generally eat leafy material, although their ability to ferment food in their hindgut allows them to subsist on more fibrous plant matter when necessary. Unlike other perissodactyls, the two African species of rhinoceros lack teeth at the front of their mouths; they rely instead on their lips to pl ...
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Tapir
Tapirs ( ) are large, herbivorous mammals belonging to the family Tapiridae. They are similar in shape to a pig, with a short, prehensile nose trunk. Tapirs inhabit jungle and forest regions of South and Central America, with one species inhabiting Southeast Asia. They are one of three extant branches of Perissodactyla (odd-toed ungulates), alongside equines and rhinoceros. Only a single genus, ''Tapirus'' is currently extant. Tapirs migrated into South America during the Pleistocene epoch from North America after the formation of the Isthmus of Panama as part of the Great American Interchange. Tapirs were once widespread in North America until the arrival of humans at the end of the Late Pleistocene, around 12,000 years ago. Species There are four widely recognized extant species of tapir, all in the genus ''Tapirus'' of the family Tapiridae. They are the South American tapir, the Malayan tapir, Baird's tapir, and the mountain tapir. In 2013, a group of researchers said they ...
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Horse
The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million years from a small multi-toed creature, ''Eohippus'', into the large, single-toed animal of today. Humans began domesticating horses around 4000 BCE, and their domestication is believed to have been widespread by 3000 BCE. Horses in the subspecies ''caballus'' are domesticated, although some domesticated populations live in the wild as feral horses. These feral populations are not true wild horses, as this term is used to describe horses that have never been domesticated. There is an extensive, specialized vocabulary used to describe equine-related concepts, covering everything from anatomy to life stages, size, colors, markings, breeds, locomotion, and behavior. Horses are adapted to run, allowing them to quickly escape predators, and po ...
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Brontothere
Brontotheriidae is a family of extinct mammals belonging to the order Perissodactyla, the order that includes horses, rhinoceroses, and tapirs. Superficially, they looked rather like rhinos, although they were actually more closely related to horses; Equidae and Brontotheriidae make up the suborder Hippomorpha. They lived around 56–34 million years ago, until the very close of the Eocene. Characteristics and evolution Brontotheres retain four toes on their front feet and three toes on their hind feet. Their teeth are adapted to shearing (cutting) relatively nonabrasive vegetation. Their molars have a characteristic W-shaped ectoloph (outer shearing blade). The evolutionary history of this group is well known due to an excellent fossil record in North America. The earliest brontotheres, such as ''Eotitanops'', were rather small, no more than a meter in height, and hornless. Brontotheres evolved massive bodies, although some small species such as ''Nanotitanops'' did pers ...
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Ungulates
Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, camels, sheep, deer, and hippopotamuses. Cetaceans such as whales, dolphins, and porpoises are also classified as even-toed ungulates, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes to support their body weight while standing or moving. The term means, roughly, "being hoofed" or "hoofed animal". As a descriptive term, "ungulate" normally excludes cetaceans as they do not possess most of the typical morphological characteristics of other ungulates, but recent discoveries indicate that they were also descended from early artiodactyls. Ungulates are typically herbivorous and many employ specialized gut bacteria to allow them to digest cellulose. Some modern species, such as pigs, are omnivorous, ...
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Perissodactyls
Odd-toed ungulates, mammals which constitute the taxonomic order Perissodactyla (, ), are animals—ungulates—who have reduced the weight-bearing toes to three (rhinoceroses and tapirs, with tapirs still using four toes on the front legs) or one (equines, third toe) of the five original toes. The non-weight-bearing toes are either present, absent, vestigial, or positioned posteriorly. By contrast, the even-toed ungulates bear most of their weight equally on four or two (an even number) of the five toes: their third and fourth toes. Another difference between the two is that odd-toed ungulates digest plant cellulose in their intestines rather than in one or more stomach chambers as even-toed ungulates, with the exception of Suina, do. The order includes about 17 species divided into three families: Equidae (horses, asses, and zebras), Rhinocerotidae (rhinoceroses), and Tapiridae (tapirs). Despite their very different appearances, they were recognized as related families in ...
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South Africa
South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countries of Namibia, Botswana, and Zimbabwe; and to the east and northeast by Mozambique and Eswatini. It also completely enclaves the country Lesotho. It is the southernmost country on the mainland of the Old World, and the second-most populous country located entirely south of the equator, after Tanzania. South Africa is a biodiversity hotspot, with unique biomes, plant and animal life. With over 60 million people, the country is the world's 24th-most populous nation and covers an area of . South Africa has three capital cities, with the executive, judicial and legislative branches of government based in Pretoria, Bloemfontein, and Cape Town respectively. The largest city is Johannesburg. About 80% of the population are Black South Afri ...
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