Amphitheriida
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Amphitheriida
Amphitheriidae is a family of Mesozoic mammals restricted to the Middle Jurassic of Britain, with indeterminate members also possibly known from the equivalently aged Itat Formation in Siberia and the Anoual Formation of Morocco. They were members of Cladotheria, more derived than members of Dryolestida, and forming a close relationship with Peramuridae.Panciroli E; Roger B.J. Benson; Richard J. Butler (2018).New partial dentaries of amphitheriid mammalian Palaeoxonodon ooliticus from Scotland, and posterior dentary morphology in early cladotherians. Acta Palaeontologica Polonica. in press. doi:10.4202/app.00434.2017. Amphitheriidae is the only family of the order Amphitheriida. Classification * Mammalia ** Yinotheria ( monotremes and relatives) ** Theriimorpha ***†Eutriconodonta ***Theriiformes ****†Allotheria *****†Gondwanatheria *****†Multituberculata **** Trechnotheria ***** † Spalacotheriidae ***** † Zhangheotheriidae ***** Cladotheria ****** † Meridiole ...
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Amphitheriida
Amphitheriidae is a family of Mesozoic mammals restricted to the Middle Jurassic of Britain, with indeterminate members also possibly known from the equivalently aged Itat Formation in Siberia and the Anoual Formation of Morocco. They were members of Cladotheria, more derived than members of Dryolestida, and forming a close relationship with Peramuridae.Panciroli E; Roger B.J. Benson; Richard J. Butler (2018).New partial dentaries of amphitheriid mammalian Palaeoxonodon ooliticus from Scotland, and posterior dentary morphology in early cladotherians. Acta Palaeontologica Polonica. in press. doi:10.4202/app.00434.2017. Amphitheriidae is the only family of the order Amphitheriida. Classification * Mammalia ** Yinotheria ( monotremes and relatives) ** Theriimorpha ***†Eutriconodonta ***Theriiformes ****†Allotheria *****†Gondwanatheria *****†Multituberculata **** Trechnotheria ***** † Spalacotheriidae ***** † Zhangheotheriidae ***** Cladotheria ****** † Meridiole ...
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Palaeoxonodon
''Palaeoxonodon'' is an extinct genus of Cladotherian mammal from the Middle Jurassic of England and Scotland. Discovery The first fossils of ''Palaeoxonodon ooliticus'' were found in the Kirtlington cement quarry, Oxfordshire, England. This site was rich in Mesozoic mammal remains from the Bathonian Forest Marble Formation. Later, two more species of ''Palaeoxonodon'' were named from the same site, ''P. leesi'' and ''P. freemani''.Sigogneau-Russell D. 2003. Holotherian mammals from the Forest Marble(Middle Jurassic of England). ''Geodiversitas'', 25, 501–537. All of these fossils were individual teeth. However, a recent fossil recovered from the Kilmaluag Formation of Skye, Scotland comprised a lower jaw with five molar teeth, four premolars, a canine and one incisor present. This more complete fossil suggests that the separate species previously named from England were in fact all the same species, ''P. ooliticus'', and only appeared different due to their differing positio ...
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Itat Formation
The Itat Formation (Russian: итатская свита) is a geologic formation in western Siberia. It was deposited in the Bajocian to Bathonian ages of the Middle Jurassic. Dinosaur remains are among the fossils that have been recovered from the formation, including the proceratosaurid ''Kileskus'', as well as fish, amphibians, mammals and many other vertebrates. The formation is noted for bearing significant coal reserves, with large open pit coal mines extracting lignite from the unit currently in operation. Lithology The lower section of the formation is around 50–130 m thick, and consists of light grey sandstones with gravel, siltstone and rare coal beds. At the Dubinino locality, the 50 m exposed section of the upper part of the formation shows an irregular rhythmic alteration of fine grained sandstone, siltstone and mudstone and coal seams typically a few tens of cm's but up to several meters thick. The deposit is located on the South Eastern margin of the West Siberi ...
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Trechnotheria
Trechnotheria is a group of mammals that includes the therians and some fossil mammals from the Mesozoic Era. In the Jurassic through Cretaceous periods, the group was endemic to what would be Asia and Africa. Trechnotheria has been assigned various ranks, but was originally described as a "superlegion" by the naming authority. A later node-based definition for Trechnotheria was the clade comprising the last common ancestor of ''Zhangheotherium'' and living therian mammals, and all its descendants. Characteristics Like most Mesozoic mammal groups, early trechnotherians are known mainly from their teeth. Hence, one of the most prominent features of this group is the "hypertrophied postvallum/prevallid shearing mechanism", along with other dental characters. Features of the shoulder blade, tibia, humerus, and ankle joint also diagnose this clade. Phylogeny A cladogram based on Luo, Cifelli & Kielan-Jaworowska, 2001, Luo, Kielan-Jaworowska & Cifelli, 2002 and, Kielan-Jaworo ...
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Eutriconodonta
Eutriconodonta is an order of early mammals. Eutriconodonts existed in Asia, Africa, Europe, North and South America during the Jurassic and the Cretaceous periods. The order was named by Kermack ''et al.'' in 1973 as a replacement name for the paraphyletic Triconodonta. Traditionally seen as the classical Mesozoic small mammalian insectivores, discoveries over the years have shown them to be among the best examples of the diversity of mammals in this time period, including a vast variety of bodyplans, ecological niches and locomotion methods. Classification "Triconodonta" had long been used as the name for an order of early mammals which were close relatives of the ancestors of all present-day mammals, characterized by molar teeth with three main cusps on a crown that were arranged in a row. The group originally included only the family Triconodontidae and taxa that were later assigned to the separate family Amphilestidae, but was later expanded to include other taxa such ...
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Anoual Formation
The Anoual Formation is a geological formation in the High Atlas of Morocco. It is early Bathonian In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. Str ... in age. It consists of two members. The lower member is several hundred metres thick, and consists largely of mudstone with lens beds of cross bedded sandstone, with thiintercalationsof limestone that was deposited in a continental setting. The upper member is several tens of metres thick and consists of limestone deposited in a shallow marine setting. The formation is fossiliferous, with several of the limestone intercalations yielding a diverse fauna, including amphibians, reptiles, dinosaurs and mammals. Paleobiota Fish Amphibians Turtles Lepidosaurs Choristoderes Dinosaurs Pterosaurs Crocodyliformes ...
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Cladotheria
Cladotheria is a clade (sometimes ranked as a legion) of mammals. It contains modern therian mammals (marsupials and placentals) and several extinct groups, such as the dryolestoids, amphitheriids and peramurids. The clade was named in 1975 by Malcolm McKenna. In 2002, it was defined as a node-based taxon containing "the common ancestor of dryolestids and living therians, plus all its descendants". A different, stem-based definition was given in 2013, in which Cladotheria contains all taxa that are closer to ''Mus musculus'' (the house mouse) than to the "symmetrodont" '' Spalacotherium tricuspidens''. Description Early cladotherians can be distinguished from other mammals by a number of derived traits (apomorphies). Their teeth differed from those of the "symmetrodonts" by the evolution of a talonid shelf ( hypoflexid) on the lower molars, which occluded with the paracone of the corresponding upper molars. A true talonid basin, allowing for the crushing and grinding of f ...
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Dryolestida
Dryolestida is an extinct order of mammals, primarily and possibly exclusively known from the Jurassic and Cretaceous. They are considered members of the clade Cladotheria, close to the ancestry of therian mammals. It is also believed that they developed a fully mammalian jaw and also had the three middle ear bones. Most members of the group, as with most Mesozoic mammals, are only known from fragmentary tooth and jaw remains. The taxonomic composition of the group is contested. Aside from the uncontroversial Dryolestidae and the possibly paraphyletic Paurodontidae, which were small insectivores, known from the Middle Jurassic to Early Cretaceous of Laurasia, the Meridiolestida, a diverse group of mammals including both small insectivores and mid-large sized herbivores known from the Late Cretaceous to Miocene of South America and possibly Antarctica, are also often included. However, in many phylogenetic analyses, Meridiolestida are recovered as an unrelated group of cladotheri ...
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Prototribosphenida
Cladotheria is a clade (sometimes ranked as a legion) of mammals. It contains modern therian mammals (marsupials and placentals) and several extinct groups, such as the dryolestoids, amphitheriids and peramurids. The clade was named in 1975 by Malcolm McKenna. In 2002, it was defined as a node-based taxon containing "the common ancestor of dryolestids and living therians, plus all its descendants". A different, stem-based definition was given in 2013, in which Cladotheria contains all taxa that are closer to ''Mus musculus'' (the house mouse) than to the " symmetrodont" '' Spalacotherium tricuspidens''. Description Early cladotherians can be distinguished from other mammals by a number of derived traits (apomorphies). Their teeth differed from those of the " symmetrodonts" by the evolution of a talonid shelf ( hypoflexid) on the lower molars, which occluded with the paracone of the corresponding upper molars. A true talonid basin, allowing for the crushing and grindin ...
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Amphitherium
''Amphitherium'' is an extinct genus of stem cladotherian mammal that lived during the Middle Jurassic of England. It was one of the first Mesozoic mammals ever described. A recent phylogenetic study found it to be the sister taxon of ''Palaeoxonodon''. It is found in the Forest Marble Formation and the Taynton Limestone Formation. Etymology ''Amphitherium'' comes from the Greek ''amphi'' meaning 'on both sides', and ''therion'' meaning 'wild beast'. This was in reference to de Blainville's incorrect belief that the original fossil jaw of this animal was not a mammal, but something in between mammals and reptiles. History The first jaws of mammals from the Mesozoic - including ''Amphitherium'' - were found in the Stonesfield Slate, part of the Taynton Limestone Formation near Stonesfield in England.Rudwick, M.J.S. 2008Worlds Before Adam/ref> These were bought by a student of the British paleontologist William Buckland. Although he thought the jaws were mammalian, the anatom ...
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Zatheria
Cladotheria is a clade (sometimes ranked as a legion) of mammals. It contains modern therian mammals (marsupials and placentals) and several extinct groups, such as the dryolestoids, amphitheriids and peramurids. The clade was named in 1975 by Malcolm McKenna. In 2002, it was defined as a node-based taxon containing "the common ancestor of dryolestids and living therians, plus all its descendants". A different, stem-based definition was given in 2013, in which Cladotheria contains all taxa that are closer to ''Mus musculus'' (the house mouse) than to the " symmetrodont" '' Spalacotherium tricuspidens''. Description Early cladotherians can be distinguished from other mammals by a number of derived traits (apomorphies). Their teeth differed from those of the " symmetrodonts" by the evolution of a talonid shelf ( hypoflexid) on the lower molars, which occluded with the paracone of the corresponding upper molars. A true talonid basin, allowing for the crushing and grinding of f ...
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Tribosphenida
Tribosphenida is a group (infralegion) of mammals that includes the ancestor of ''Hypomylos'', Aegialodontia and Theria (the last common ancestor of marsupials and placentals plus all of its descendants). Its current definition is more or less synonymous with Boreosphenida. Characteristics Tribosphenid mammals were originally grouped on the basis of triangular or V-shaped ( tribosphenic) molars. Since then, other unrelated mammal groups have been found to have tribosphenic molars, such as the australosphenidans (a group that includes the still extant monotremes), suggesting that as a synapomorphy this is fundamentally useless as it evolved multiple times among mammals. However, a clade between the aforementioned groups, the "true Tribosphenida" or Boreosphenida, is still identifiable, united by characteristics such as the lack of a mesial cingulid and of a triangulated trigonid on the last premolar. They are also united by postcranial features such as the presence of a modern ea ...
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