Allodaposuchidae
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Allodaposuchidae
Allodaposuchidae is an extinct clade of eusuchians that lived in Europe during the Late Cretaceous (Santonian-Maastrichtian). Systematics The type genus, ''Allodaposuchus'', was originally described in 1928 by Nopcsa from the Maastrichtian-age Sard Formation of the Hațeg Basin in Transylvania, Romania, and classified as a relative of the North American ''Leidyosuchus''. It was later classified as a eusuchian outside of Crocodylia in a 2001 paper, and subsequent studies found a number of European eusuchian species ('' Arenysuchus'', '' Ischyrochampsa'', '' Massaliasuchus'', ''Musturzabalsuchus'') to group with ''Allodaposuchus'', prompting the erection of the clade Allodaposuchidae to accommodate ''Allodaposuchus'' and all European eusuchians closely related to it. Narváez ''et al.'' cladistically defined Allodaposuchidae in 2015 as '' Allodaposuchus precedens'' and all crocodyliforms more closely related to it than to ''Hylaeochampsa vectiana'', '' Shamosuchus djadochtaensis' ...
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Allodaposuchus Precedens
''Allodaposuchus'' is an extinct genus of crocodyliforms that lived in what is now southern Europe during the Campanian and Maastrichtian stages of the Late Cretaceous. Although generally classified as a non-crocodylian eusuchian crocodylomorph, it is sometimes placed as one of the earliest true crocodylians. ''Allodaposuchus'' is one of the most common Late Cretaceous crocodylomorphs from Europe, with fossils known from Romania, Spain, and France. Description Like many other Cretaceous crocodylomorphs, ''Allodaposuchus'' has a relatively small body size compared to living crocodylians. The largest known specimen of ''Allodaposuchus'' belongs to an individual that was probably around long. Although the shape varies between species, in general ''Allodaposuchus'' has a short, flattened, and rounded skull. ''Allodaposuchus precedens'' has a brevirostrine or "short-snouted" skull with a snout about the same length as the skull table (the region of the skull behind the eye sockets) a ...
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Allodaposuchus
''Allodaposuchus'' is an extinct genus of crocodyliforms that lived in what is now southern Europe during the Campanian and Maastrichtian stages of the Late Cretaceous. Although generally classified as a non-crocodylian eusuchian crocodylomorph, it is sometimes placed as one of the earliest true crocodylians. ''Allodaposuchus'' is one of the most common Late Cretaceous crocodylomorphs from Europe, with fossils known from Romania, Spain, and France. Description Like many other Cretaceous crocodylomorphs, ''Allodaposuchus'' has a relatively small body size compared to living crocodylians. The largest known specimen of ''Allodaposuchus'' belongs to an individual that was probably around long. Although the shape varies between species, in general ''Allodaposuchus'' has a short, flattened, and rounded skull. ''Allodaposuchus precedens'' has a brevirostrine or "short-snouted" skull with a snout about the same length as the skull table (the region of the skull behind the eye sockets) a ...
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Lohuecosuchus
''Lohuecosuchus'' (meaning "Lo Hueco crocodile") is an extinct genus of Allodaposuchidae, allodaposuchid eusuchian Crocodylomorpha, crocodylomorph that lived during the Late Cretaceous (late Campanian to early Maastrichtian) in what is now Spain and southern France. Description The anatomy of the endocranium (braincase) of ''L. megadontos'' was found to be similar to crown-crocodylians, revealing that the acute sense of olfaction and low frequency hearing found in living crocodylians likely originated along the stem-line. Classification ''Lohuecosuchus'' is a member of the clade Allodaposuchidae, a group of eusuchians that lived in southern Europe during the Late Cretaceous. Two species of ''Lohuecosuchus'' were scientifically described, described in 2015. The type species, ''Lohuecosuchus megadontos'', was recovered from the Lo Hueco fossil site of the Villalba de la Sierra Formation in the municipality of Fuentes, Cuenca, Fuentes, Cuenca Province at Castilla-La Mancha of centr ...
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Eusuchia
Eusuchia is a clade of crocodylomorphs that first appeared in the Early Cretaceous with ''Hylaeochampsa''. Along with Dyrosauridae and Sebecosuchia, they were the only crocodyliformes who survived the K-T extinction. Since the other two clades died out 47 and 11 million years ago respectively, all living crocodilian species are eusuchians, as are many extinct forms. Definition Eusuchia was originally defined by Thomas Henry Huxley in 1875 as an apomorphy-based group, meaning that it was defined by shared characteristics rather than relations. These characteristics include pterygoid-bounded choanae and vertebrae which are procoelous (concave from the front and convex from the back). The possibility that these traits may have been convergently evolved in different groups of neosuchians rather than one lineage spurred some modern paleontologists to revise the group's definition to make it defined solely by relations. In 1999, Christopher Brochu redefined Eusuchia as "the last common ...
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Musturzabalsuchus
''Musturzabalsuchus'' is an extinct monospecific genus of allodaposuchid eusuchian crocodyliform. The type and only species is ''Musturzabalsuchus buffetauti''. Etymology The generic name means "broadened rostrum crocodile", with "Musturzabal" meaning "broadened rostrum" in Basque and "suchus" meaning "crocodile" in Greek. The type and only species is ''M. buffetauti'', named after the French paleoherpetologist Eric Buffetaut. Discovery The material first assigned to ''Musturzabalsuchus'' in 1997 has been found from the locality of Laño in Condado de Treviño, northern Spain. Although dating back to the Late Cretaceous, the exact age of the strata in which material of ''Musturzabalsuchus'' occurs in the locality is not known: it is either Late Campanian or very Early Maastrichtian. Despite the unusually high quantity of remains belonging to the genus, the only skeletal elements known from ''Musturzabalsuchus'' are the maxilla and mandible. Some fragments of these bones have ...
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Planocrania Datangensis
''Planocrania'' is an extinct genus of eusuchian crocodyliforms from what is now China. Two species are currently known to belong to the genus. History The type species, ''Planocrania datangensis'', was named in 1976 from material found from Nanxiong in Guangdong Province, China, and the new genus ''Planocrania'' was placed in a newly erected family Planocraniidae. A second species, ''Planocrania hengdongensis'', is known from Hengdong County in Hunan Province and was described in 1984. ''Planocrania'' was subsequently included in the family Pristichampsidae along with the genus ''Pristichampsus''. Both genera were previously assigned to the subfamily Pristichampsinae within the family Crocodylidae before they were placed in their own family, the Pristichampsidae. However, in a 2013 phylogenetic analysis, the two Asian species of ''Planocrania'' were found to be most closely related to the North American and European '' Boverisuchus'', and the family Planocraniidae was reinsta ...
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Arenysuchus
''Arenysuchus'' (meaning "Arén crocodile") is an extinct monospecific genus of allodaposuchid eusuchian crocodylomorph from Late Cretaceous (late Maastrichtian stage) deposits of north Spain. It is known from the holotype MPZ ELI-1, a partial skull from Elías site, and from the referred material MPZ2010/948, MPZ2010/949, MPZ2010/950 and MPZ2010/951, four teeth from Blasi 2 site. It was found by the researchers José Manuel Gasca and Ainara Badiola from the Tremp Formation, in Arén of Huesca, Spain. It was first named by Eduardo Puértolas, José I. Canudo and Penélope Cruzado-Caballero in 2011 and the type species is ''Arenysuchus gascabadiolorum''. Etymology ''Arenysuchus'' was named and described in 2011 by Eduardo Puértolas and his colleagues for a partial skull and teeth. For the generic name, Areny is named after Arén, spelt as Areny in the Catalan language, the locality where the skull was found, and ''souchus'', the Greek word for crocodile, leading ...
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Massaliasuchus
''Massaliasuchus'' is an extinct monospecific genus of allodaposuchid eusuchian crocodyliform that is known from fossils found in Santonian–Campanian-age Upper Cretaceous rocks of southeastern France. Systematics ''Massaliasuchus'' was first described in 1869 by Philippe Matheron as '' Crocodilus affuvelensis'', based on remains including skull bones. The new genus name was given to it in 2008 by Jeremy Martin and Eric Buffetaut. ''Massaliasuchus'' was considered to be related to early alligatoroids. Its name means "Marseille crocodile". Recent cladistic analysis places ''Massaliasuchus'' as a member of Allodaposuchidae, a clade of basal eusuchians from the Late Cretaceous of southern Europe. The genus can be distinguished from ''Musturzabalsuchus'' in having 15 alveoli in the dentary. However, ''Massaliasuchus'' is usually omitted from phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενε ...
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Agaresuchus
''Agaresuchus'' is an extinct genus of allodaposuchid eusuchian crocodylomorph from the Late Cretaceous (Campanian-Maastrichtian) of Spain. It includes two species, the type species ''Agaresuchus fontisensis'', and ''Agaresuchus subjuniperus'', which was originally named as a species of the related genus ''Allodaposuchus''. However, it has been proposed that both species may instead belong to the genus ''Allodaposuchus''. Discovery and naming The genus ''Agaresuchus'' was named in 2016 upon the discovery of '' Agaresuchus fontisensis''. ''Allodaposuchus subjuniperus'', discovered in 2013 and originally classified as a new and second species of ''Allodaposuchus'', was then reassigned to ''Agaresuchus''. ''A. subjuniperus'' was named in 2013 on the basis of a skull from the late Maastrichtian-aged Conquès Formation, part of the Tremp Group, in the province of Huesca, Spain. The skull was found underneath a juniper tree whose roots had grown between the bones, hence the species n ...
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Ischyrochampsa
''Ischyrochampsa'' is an extinct monospecific genus of Late Cretaceous crocodyliform belonging to the eusuchian clade Allodaposuchidae. Fossils of the type species ''I. meridionalis'' are late Campanian in age and were found in the commune of Saint-Estève-Janson in Bouches-du-Rhône, France. Material is also known from Spain. It was named and scientifically described, described in 1995, and it had an estimated length of over . ''Ischyrochampsa'' was first classified as a trematochampsid, but was removed from the group by subsequent studies. In their description of ''Allodaposuchus'' remains from southern France, Martin and his colleagues treated the genus as a possible junior synonym of ''Allodaposuchus''.Martin, J. E., Delfino, M., Garcia, G., Godefroit, P., Berton, S. and Valentin, X. (2016), New specimens of Allodaposuchus precedens from France: intraspecific variability and the diversity of European Late Cretaceous eusuchians. Zoological Journal of the Linnean Society, 176: 6 ...
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Borealosuchus Sternbergii
''Borealosuchus'' (meaning "Subarctic climate, boreal crocodile") is an extinct genus of crocodyliforms that lived from the Late Cretaceous to the Eocene in North America. It was named by Chris Brochu in 1997 for several species that had been assigned to ''Leidyosuchus''. The species assigned to it are: ''B. sternbergii'', the type species, from the Maastrichtian (Late Cretaceous) of Colorado, Montana, North Dakota, South Dakota, and Wyoming; ''B. acutidentatus'', from the Paleocene of Saskatchewan; ''B. formidabilis'', from the Paleocene of North Dakota; ''B. griffithi'', from the Paleocene of Alberta; and ''B. wilsoni'', from the Eocene of Wyoming. ''B. formidabilis'' is particularly well-known, represented by the remains of many individuals from the Wannagan Creek site in North Dakota. ''Borealosuchus'' was a mid-sized crocodyliform; ''B. acutidentatus'' reached up to in length with a skull. Taxonomy Six species of ''Borealosuchus'' are currently recognized. In order of th ...
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Cladistic
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived characteristics ( synapomorphies'')'' that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms ''worms'' or ''fishes'' were used within a ''strict'' cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically, outside of cladistics, e.g. as a 'grade', which are fruitless to precisely delineate, especially when including extinct species. R ...
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