Alcione (pterosaur)
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Alcione (pterosaur)
''Alcione'' is a genus of nyctosaurid pterosaur from the Ouled Abdoun Basin of Morocco, dating back to the Maastrichtian stage of the Late Cretaceous period. Only one species, ''A. elainus'', is known. This pterosaur lived in a marine environment alongside several other pterosaurs, including the nyctosaurids ''Simurghia'' and ''Barbaridactylus''. Discovery and naming All known specimens of ''Alcione'' were uncovered in a 3-year dig that unearthed about 200 pterosaur specimens. The type specimen, FSAC-OB 2, consists of a partial skeleton that preserves various elements, including the femur, humerus, scapulocoracoid, and sternum. Referred specimens include FSAC-OB 217, which consists of metacarpal IV, FSAC-OB 156, a mandible, and the more complete FSAC-OB 4. This specimen consists of the humerus, radius, ulna, a partial metacarpal IV, phalanx IV-1, scapulocoracoids, and synsacra. ''Alcione'' is named after Alcyone, who was transformed into a seabird, from Greek mythology. The ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Ornithocheiroidea
Ornithocheiroidea (or ornithocheiroids) is a group of pterosaurs within the extinct suborder Pterodactyloidea. They were typically large pterosaurs that lived from the Early to Late Cretaceous periods (Valanginian to Maastrichtian stages), with fossil remains found all over the world except Antarctica. Ornithocheiroids were the most advanced group of pterosaurs, as the group includes the clade Azhdarchoidea, of which its members lived until the Maastrichtian stage of the Late Cretaceous, around 66 million years ago. Notable pterosaurs from this group include the pteranodontians ''Pteranodon'' and ''Nyctosaurus'', the ornithocheirid ''Ornithocheirus'', the anhanguerid ''Tropeognathus'', as well as the azhdarchids ''Hatzegopteryx'' and ''Quetzalcoatlus''. Classification The name Ornithocheiroidea was originally defined as an apomorphy-based taxon by Christopher Bennett in 1994. It was given a relationship-based definition in 2003 by Alexander Kellner, who defined it as the least in ...
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Simurghia Robusta
''Simurghia'' is a genus nyctosaurid pterosaur from the Ouled Abdoun Basin of Morocco, a basin that dates to the Maastrichtian stage of the Late Cretaceous period, about 66 million years ago. It was published in 2018 by paleontologists Nicholas R. Longrich, David M. Martill, and Brian Andres, along with two other nyctosaurids from the same basin: ''Alcione'' and ''Barbaridactylus''. The type and only species is ''S. robusta''. Discovery and naming All known specimens of ''Simurghia'' were uncovered in a 3-year dig that unearthed about 200 pterosaur specimens. The type specimen, FSAC-OB 7, consists of a nearly complete humerus, lacking only the humeral head and the ulnar crest. ''Simurghia'' is named after a flying beast from Persian mythology known as the Simurgh. The species name, ''S. robusta'', is a Latin word that means "robust." Classification Below is a cladogram showing the results of a phylogenetic analysis first presented by Andres and colleagues in 2014, and updated ...
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Cretornis Hlavaci
''Cretornis'' is a pterosaur genus from the late Cretaceous period (Turonian stage) of what is now the Jizera Formation in the Czech Republic, dating to about 92 million years ago. It only contains a single species, ''Cretornis hlavaci''. Discovery and naming The fossils were discovered in 1880 by workers at a quarry in Zářecká Lhota near the town of Choceň, who were getting gravel to repair a local road. A certain Mrs. Tomková, a croupier from Choceň, then alerted František Hlaváč, a Choceň pharmacist and fossil collector, to the find. He recognised exceptionality of that one, secured the rest of the fossils and then sent them to naturalist Professor Antonín Frič in Prague. In 1881, Antonín Frič identified it as a pterosaur and named it as the type species ''Cretornis Hlaváči''. The generic name is derived from Latin ''creta'', "chalk", in reference to the Cretaceous, and Greek ὄρνις, ''ornis'', "bird", as Frič originally thought that the fossil bones ...
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Volgadraco Bogolubovi
''Volgadraco'' ("Volga River dragon") is a genus of pterodactyloid pterosaur from the Upper Cretaceous of European Russia. ''Volgadraco'' was originally classified as an azhdarchid. However, recent studies have concluded that it may belong to either the family Nyctosauridae, or the family Pteranodontidae. ''Volgadraco'' is known from lower beak (holotype SGU, no. 46/104a) and postcranial fragments from the early Campanian-age Rybushka Formation of Saratov, Russia. The size of this animal, and the development of blood supply in the lower jaw are intermediate between older Santonian or Turonian azhdarchids like ''Azhdarcho'' and '' Bakonydraco'' and later Maastrichtian azhdarchids like ''Quetzalcoatlus''. ''Volgadraco'' was described in 2008 by Averianov, Arkhangelsky, and Pervushov. The type species is ''V. bogolubovi'', the specific name honouring Russian paleontologist Nikolai Nikolaevich Bogolubov. The authors consider the earlier named genus ''Bogolubovia'' to be a ''nomen ...
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Alamodactylus Byrdi
''Alamodactylus'' is an extinct genus of nyctosaurid pteranodontoid pterosaur known from the Late Cretaceous of Texas, southern United States. It contains a single species, ''Alamodactylus byrdi''. Discovery ''Alamodactylus'' is known solely from its holotype, SMU 76476, a partial left wing first described by Myers (2010).Myers, T.S., 2010, "Earliest occurrence of the Pteranodontidae (Archosauria: Pterosauria) in North America: New material from the Austin Group of Texas", ''Journal of Paleontology'' 84(6): 1071–1081 This specimen is crushed and consists of a left humerus, the distal end of the fourth wing metacarpal and the proximal end of the first wing phalanx of fourth digit. Other elements of the holotype first reported by Andres and Myers (2013) include a left proximal syncarpal, manual phalanx, and some fragments including four tapering processes that may represent skull bones, an articular surface that appears to be the mandible articulation of the right quadrate ...
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Tethydraco Regalis
''Tethydraco'' is a genus of pterodactyloid pterosaur that lived during the Late Cretaceous period (Maastrichtian stage) of what is now the area of present Morocco, about 66 million years ago. ''Tethydraco'' was originally assigned to the family Pteranodontidae. Subsequently described material suggested that it was an azhdarchid, and possibly synonymous with ''Phosphatodraco''. The type and only species is ''T. regalis''. Discovery and naming Since 2015, a group of paleontologists has been acquiring pterosaur fossils from commercial Moroccan fossil traders, who obtain these from workers in the phosphate mines on the Khouribga plateau, which is located within the Ouled Abdoun Basin. The purpose of this project is to determine pterosaur diversity in the latest Cretaceous. From this stage, no Konservat-Lagerstätten are known, sites combining a large variety of species with exceptional preservation. It is in such sites that the vast majority of pterosaur fossils and taxa have bee ...
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Pteranodon Longiceps
''Pteranodon'' (); from Ancient Greek (''pteron'', "wing") and (''anodon'', "toothless") is a genus of pterosaur that included some of the largest known flying reptiles, with ''P. longiceps'' having a wingspan of . They lived during the late Cretaceous geological period of North America in present-day Kansas, Nebraska, Wyoming, South Dakota and Alabama. More fossil specimens of ''Pteranodon'' have been found than any other pterosaur, with about 1,200 specimens known to science, many of them well preserved with nearly complete skulls and articulated skeletons. It was an important part of the animal community in the Western Interior Seaway. ''Pteranodon'' was not a dinosaur. By definition, all dinosaurs belong to the group Dinosauria; ''Pteranodon'' belongs to the group Pterosauria. Nonetheless, ''Pteranodon'' is the most famous pterosaur, frequently featured in dinosaur media and strongly associated with dinosaurs by the general public. While not dinosaurs, pterosaurs such as ' ...
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Pteranodon Sternbergi
''Geosternbergia'' is an extinct genus of pteranodontid pterodactyloid pterosaur from the Late Cretaceous geological period of North America. ''Geosternbergia'' was one of the largest pterosaur genera, and had a wingspan of up to . Discovery and history The first fossils of ''Geosternbergia'' were collected by American paleontologist George F. Sternberg in 1952 from the lower portion of the Niobrara Formation. The fossils of the animal looked similar to those of the species ''Pteranodon longiceps'', but the crests were set upright and in a slightly different position. In 1958, Sternberg and paleontologist Myrl V. Walker published a study about this peculiar find. In 1966, American paleontologist John Christian Harksen assigned the specimens found as a new species of ''Pteranodon'' called ''P. sternbergi'' due to its distinct upright crest that set it apart from ''P. longiceps''. Halsey W. Miller however, concluded a revision of the different species of ''Pteranodon'' in 197 ...
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Pteranodontidae
The Pteranodontidae are a family of large pterosaurs of the Cretaceous Period of North America and Africa. The family was named in 1876 by Othniel Charles Marsh. Pteranodontids had a distinctive, elongated crest jutting from the rear of the head (most famously seen in ''Pteranodon'' itself). The spectacularly-crested ''Nyctosaurus'' is sometimes included in this family, though usually placed in its own family, the Nyctosauridae (Nicholson & Lydekker, 1889). Modern researchers differ in their use of the concept. S. Christopher Bennett and Alexander Kellner have concluded that ''Nyctosaurus'' was not a pteranodontid. In 1994 Bennett defined a clade Pteranodontidae, also including species of the Anhangueridae. However, this definition has not been accepted by other workers. Alexander Kellner, for example, named several additional species for specimens previously classified as ''Pteranodon'', and placed ''P. sternbergi'' in a distinct genus, ''Geosternbergia''. Kellner re-defined Ptera ...
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Pteranodontia
Pteranodontia is an extinct group of ornithocheiroid pterodactyloid pterosaurs that lived during the Late Cretaceous period (Coniacian to Maastrichtian stages) of North America and Africa. They were some of the most advanced pterosaurs, and possessed highly specialized cranial crests that may have served as sexual attraction, with males having a much larger crest. Classification Pteranodontia was originally named by Marsh in 1876. In 2003, it was given a phylogenetic definition by David Unwin as the common ancestor of ''Pteranodon'' and ''Nyctosaurus'' plus all its descendants. Though Marsh had originally named this group based on the shared absence of teeth in those species, most analyses show that all of the traditional "ornithocheiroid" pterosaurs are also members of this clade. Below is a cladogram showing the phylogenetic placement of this group from Andres and Myers (2013). In 2018, Longrich, Martill, and Andres revisited the classification, and made a different phylogen ...
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Ornithocheiromorpha
Ornithocheiromorpha (from Ancient Greek, meaning "bird hand form") is a group of pterosaurs within the suborder Pterodactyloidea. Fossil remains of this group date back from the Early Cretaceous, Early to Late Cretaceous Period (geology), periods (Valanginian to Turonian stages), around 140 to 92.5 million years ago. Ornithocheiromorphs were discovered worldwide except Antarctica, though most genera were recovered in Europe, Asia and South America. They were the most diverse and successful pterosaurs during the Early Cretaceous, but throughout the Late Cretaceous they were replaced by better adapted and more advanced pterosaur species such the pteranodontids and azhdarchoids. The Ornithocheiromorpha was defined in 2014 by Andres and colleagues, and they made Ornithocheiromorpha the most inclusive clade containing ''Ornithocheirus'', but not ''Pteranodon''. Ornithocheiromorphs are considered to be some of the largest animals to have ever flown. Members of this group are also rega ...
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