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Adaptationism
Adaptationism (also known as functionalism) is the Darwinian view that many physical and psychological traits of organisms are evolved adaptations. Pan-adaptationism is the strong form of this, deriving from the early 20th century modern synthesis, that all traits are adaptations, a view now shared by only a few biologists. The "adaptationist program" was heavily criticized by Stephen Jay Gould and Richard Lewontin in their 1979 paper "The Spandrels of San Marco and the Panglossian Paradigm". According to Gould and Lewontin, evolutionary biologists had a habit of proposing adaptive explanations for any trait by default without considering non-adaptive alternatives, and often by conflating products of adaptation with the process of natural selection. One formal alternative to adaptationist explanations for traits in organisms is the neutral theory of molecular evolution, which proposes that features in organisms can arise through neutral transitions and become fixed in a population ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
The Spandrels Of San Marco And The Panglossian Paradigm
"The Spandrels of San Marco and the Panglossian Paradigm: A Critique of the Adaptationist Programme", also known as the "Spandrels paper", is a paper by evolutionary biologists Stephen Jay Gould and Richard Lewontin, originally published in the ''Proceedings of the Royal Society B: Biological Sciences'' in 1979. The paper criticizes the adaptationist school of thought that was prevalent in evolutionary biology at the time using two metaphors: that of the spandrels in St Mark's Basilica, a cathedral in Venice, Italy, and that of the fictional character "Pangloss" in Voltaire's novella ''Candide''. The paper was the first to use the architecture, architectural term "spandrel" in a biology, biological context; the term "spandrel (biology), spandrel" has since gained currency in biology to refer to byproducts of adaptation. Background "Spandrels" was originally written in 1978, and that year Gould delivered it as a talk to the Royal Society. Gould had visited St. Mark's Cathedral short ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Constructive Neutral Evolution
Constructive neutral evolution (CNE) is a theory that seeks to explain how complex systems can evolve through neutral transitions and spread through a population by chance fixation (genetic drift). Constructive neutral evolution is a competitor for both adaptationist explanations for the emergence of complex traits and hypotheses positing that a complex trait emerged as a response to a deleterious development in an organism. Constructive neutral evolution often leads to irreversible or "irremediable" complexity and produces systems which, instead of being finely adapted for performing a task, represent an excess complexity that has been described with terms such as "runaway bureaucracy" or even a "Rube Goldberg machine". The groundworks for the concept of CNE were laid by two papers in the 1990s, although first explicitly proposed by Arlin Stoltzfus in 1999. The first proposals for the role CNE was in the evolutionary origins of complex macromolecular machines such as the spliceosom ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Stephen Jay Gould
Stephen Jay Gould (; September 10, 1941 – May 20, 2002) was an American paleontologist, evolutionary biologist, and historian of science. He was one of the most influential and widely read authors of popular science of his generation. Gould spent most of his career teaching at Harvard University and working at the American Museum of Natural History in New York. In 1996, Gould was hired as the Vincent Astor Visiting Research Professor of Biology at New York University, after which he divided his time teaching between there and Harvard. Gould's most significant contribution to evolutionary biology was the theory of punctuated equilibrium developed with Niles Eldredge in 1972.Eldredge, Niles, and S. J. Gould (1972)"Punctuated equilibria: an alternative to phyletic gradualism."In T.J.M. Schopf, ed., ''Models in Paleobiology''. San Francisco: Freeman, Cooper and Company, pp. 82–115. The theory proposes that most evolution is characterized by long periods of evolutionary stability, ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Richard Lewontin
Richard Charles Lewontin (March 29, 1929 – July 4, 2021) was an American evolutionary biologist, mathematician, geneticist, and social commentator. A leader in developing the mathematical basis of population genetics and evolutionary theory, he pioneered the application of techniques from molecular biology, such as gel electrophoresis, to questions of genetic variation and evolution. In a pair of seminal 1966 papers co-authored with J. L. Hubby in the journal ''Genetics'', Lewontin helped set the stage for the modern field of molecular evolution. In 1979 he and Stephen Jay Gould introduced the term "spandrel" into evolutionary theory. From 1973 to 1998, he held an endowed chair in zoology and biology at Harvard University, and from 2003 until his death in 2021 he was a research professor there. Lewontin opposed genetic determinism. Early life and education Lewontin was born in New York City, to parents descended from late 19th-century Ashkenazi Jewish immigrants. His father ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Neutral Theory Of Molecular Evolution
The neutral theory of molecular evolution holds that most evolutionary changes occur at the molecular level, and most of the variation within and between species are due to random genetic drift of mutant alleles that are selectively neutral. The theory applies only for evolution at the molecular level, and is compatible with phenotypic evolution being shaped by natural selection as postulated by Charles Darwin. The neutral theory allows for the possibility that most mutations are deleterious, but holds that because these are rapidly removed by natural selection, they do not make significant contributions to variation within and between species at the molecular level. A neutral mutation is one that does not affect an organism's ability to survive and reproduce. The neutral theory assumes that most mutations that are not deleterious are neutral rather than beneficial. Because only a fraction of gametes are sampled in each generation of a species, the neutral theory suggests that a mu ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Evolution Of The Eye
Many scientists have found the evolution of the eye attractive to study because the eye distinctively exemplifies an analogous organ found in many animal forms. Simple light detection is found in bacteria, single-celled organisms, plants and animals. Complex, image-forming eyes have evolved independently several times. Diverse eyes are known from the Burgess shale of the Middle Cambrian, and from the slightly older Emu Bay Shale. Eyes vary in their visual acuity, the range of wavelengths they can detect, their sensitivity in low light, their ability to detect motion or to resolve objects, and whether they can discriminate colours. History of research In 1802, philosopher William Paley called it a miracle of "design." In 1859, Charles Darwin himself wrote in his ''Origin of Species'', that the evolution of the eye by natural selection seemed at first glance "absurd in the highest possible degree". However, he went on that despite the difficulty in imagining it, its evolut ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Randolph Nesse
Randolph M. Nesse (born 1948) is an American physician, scientist and author who is notable for his role as a founder of the field of evolutionary medicine and evolutionary psychiatry. He is professor of life sciences and ASU Foundation Professor at Arizona State University, where he became the Founding Director of the Center for Evolution and Medicine in 2014. He was previously a professor of psychiatry, professor of psychology and research professor at the University of Michigan where he led the Evolution and Human Adaptation Program and helped to establish one of the world’s first anxiety Anxiety is an emotion which is characterized by an unpleasant state of inner turmoil and includes feelings of dread over anticipated events. Anxiety is different than fear in that the former is defined as the anticipation of a future threat wh ... disorders clinics and conducted research on neuroendocrine responses to fear. His research on the evolution of aging led to a long collabor ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Blind Spot (vision)
A blind spot, scotoma, is an obscuration of the visual field. A particular blind spot known as the ''physiological blind spot'', "blind point", or ''punctum caecum'' in medical literature, is the place in the visual field that corresponds to the lack of light-detecting photoreceptor cells on the optic disc of the retina where the optic nerve passes through the optic disc.Gregory, R., & Cavanagh, P. (2011)"The Blind Spot" Scholarpedia. Retrieved on 2011-05-21. Because there are no cells to detect light on the optic disc, the corresponding part of the field of vision is invisible. Processes in the brain interpolate the blind spot based on surrounding detail and information from the other eye, so it is not normally perceived. Although all vertebrates have this blind spot, cephalopod eyes, which are only superficially similar, do not. In them, the optic nerve approaches the receptors from behind, so it does not create a break in the retina. The first documented observation of the ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Retina
The retina (from la, rete "net") is the innermost, light-sensitive layer of tissue of the eye of most vertebrates and some molluscs. The optics of the eye create a focused two-dimensional image of the visual world on the retina, which then processes that image within the retina and sends nerve impulses along the optic nerve to the visual cortex to create visual perception. The retina serves a function which is in many ways analogous to that of the film or image sensor in a camera. The neural retina consists of several layers of neurons interconnected by synapses and is supported by an outer layer of pigmented epithelial cells. The primary light-sensing cells in the retina are the photoreceptor cells, which are of two types: rods and cones. Rods function mainly in dim light and provide monochromatic vision. Cones function in well-lit conditions and are responsible for the perception of colour through the use of a range of opsins, as well as high-acuity vision used for task ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Cephalopod
A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by ''Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been ident ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Darwinism
Darwinism is a scientific theory, theory of Biology, biological evolution developed by the English naturalist Charles Darwin (1809–1882) and others, stating that all species of organisms arise and develop through the natural selection of small, inherited variations that increase the individual's ability to compete, survive, and reproduction, reproduce. Also called Darwinian theory, it originally included the broad concepts of transmutation of species or of evolution which gained general scientific acceptance after Darwin published ''On the Origin of Species'' in 1859, including concepts which predated Darwin's theories. English biologist Thomas Henry Huxley coined the term ''Darwinism'' in April 1860. Terminology Darwinism subsequently referred to the specific concepts of natural selection, the Weismann barrier, or the central dogma of molecular biology. Though the term usually refers strictly to biological evolution, creationism, creationists have appropriated it to refer to ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
Cranial Nerves
Cranial nerves are the nerves that emerge directly from the brain (including the brainstem), of which there are conventionally considered twelve pairs. Cranial nerves relay information between the brain and parts of the body, primarily to and from regions of the head and neck, including the special senses of vision, taste, smell, and hearing. The cranial nerves emerge from the central nervous system above the level of the first vertebra of the vertebral column. Each cranial nerve is paired and is present on both sides. There are conventionally twelve pairs of cranial nerves, which are described with Roman numerals I–XII. Some considered there to be thirteen pairs of cranial nerves, including cranial nerve zero. The numbering of the cranial nerves is based on the order in which they emerge from the brain and brainstem, from front to back. The terminal nerves (0), olfactory nerves (I) and optic nerves (II) emerge from the cerebrum, and the remaining ten pairs arise from the ... [...More Info...] [...Related Items...] OR: [Wikipedia] [Google] [Baidu] |
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