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Azendohsaurus Madagaskarensis
''Azendohsaurus'' is an extinct genus of herbivorous archosauromorph reptile from roughly the late Middle to early Late Triassic Period of Morocco and Madagascar. The type species, ''Azendohsaurus laaroussii'', was described and named by Jean-Michel Dutuit in 1972 based on partial jaw fragments and some teeth from Morocco. A second species from Madagascar, ''A. madagaskarensis'', was first described in 2010 by John J. Flynn and colleagues from a multitude of specimens representing almost the entire skeleton. The generic name "Azendoh lizard" is for the village of Azendoh, a local village near where it was first discovered in the Atlas Mountains. It was a bulky quadruped that unlike other early archosauromorphs had a relatively short tail and robust limbs that were held in an odd mix of sprawled hind limbs and raised forelimbs. It had a long neck and a proportionately small head with remarkably sauropod-like jaws and teeth. ''Azendohsaurus'' used to be classified as a herbivo ...
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Middle Triassic
In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between Ma and Ma (million years ago). It is preceded by the Early Triassic Epoch and followed by the Late Triassic Epoch. The Middle Triassic is divided into the Anisian and Ladinian ages or stages. Formerly the middle series in the Triassic was also known as Muschelkalk. This name is now only used for a specific unit of rock strata with approximately Middle Triassic age, found in western Europe. Middle Triassic fauna Following the Permian–Triassic extinction event, the most devastating of all mass-extinctions, life recovered slowly. In the Middle Triassic, many groups of organisms reached higher diversity again, such as the marine reptiles (e.g. ichthyosaurs, sauropterygians, thallatosaurs), ray-finned fish and many invertebrate groups like ...
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Atlas Mountains
The Atlas Mountains are a mountain range in the Maghreb in North Africa. It separates the Sahara Desert from the Mediterranean Sea and the Atlantic Ocean; the name "Atlantic" is derived from the mountain range. It stretches around through Morocco, Algeria and Tunisia. The range's highest peak is Toubkal, which is in central Morocco, with an elevation of . The Atlas mountains are primarily inhabited by Berbers, Berber populations. The terms for 'mountain' are ''Adrar'' and ''adras'' in some Berber languages, and these terms are believed to be cognates of the Toponymy, toponym ''Atlas''. The mountains are also home to a number of animals and plants which are mostly found within Africa but some of which can be found in Europe. Many of these species are endangered and a few are already extinct. The weather is cooling but has sunny summers, and the average temperature there is 25°C.Atlas Mountains are a mountain range in the Maghreb in North Africa. It separates the Sahara Desert fro ...
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Shringasaurus
''Shringasaurus'' (meaning "horned lizard", from Sanskrit शृङ्ग (''śṛṅga),'' "horn", and Ancient Greek (''sauros),'' "lizard") is an extinct genus of archosauromorph reptile from the Middle Triassic (Anisian) of India. It is known from the type and only known species, ''S. indicus''. ''Shringasaurus'' is known from the Denwa Formation in the state of Madhya Pradesh. ''Shringasaurus'' was an allokotosaur, a group of unusual herbivorous reptiles from the Triassic, and is most closely related to the smaller and better known ''Azendohsaurus'' in the family Azendohsauridae. Like some ceratopsid dinosaurs, ''Shringasaurus'' had two large horns over its eyes that faced up and forwards from its skull. These horns were likely used for display, and possibly during fights with other ''Shringasaurus'', much like what has been suggested for the horns of ceratopsids like ''Triceratops''. ''Shringasaurus'' also bears similarities to sauropodomorph dinosaurs, such as its long nec ...
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Azendohsauridae
Azendohsauridae is a family of allokotosaurian archosauromorphs that lived during the Middle to Late Triassic period, around 242-216 million years ago. The family was originally named solely for the eponymous ''Azendohsaurus'', marking out its distinctiveness from other allokotosaurs, but the family now includes four other genera: the basal genus '' Pamelaria'', the large horned herbivore '' Shringasaurus'', and two carnivorous genera grouped into the subfamily-level subclade Malerisaurinae, '' Malerisaurus'' and '' Puercosuchus'', and potentially also the dubious genus ''Otischalkia''. Most fossils of azendohsaurids have a Gondwanan distribution, with multiple species known across Morocco and Madagascar in Africa as well as India, although fossils of malerisaurine azendohsaurids have also been found in the southwestern United States of North America. Azendohsaurids are notable for the various dinosaur-like traits found in some species, including the sauropodomorph-like nec ...
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Family (biology)
Family ( la, familia, plural ') is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as the "walnut family". What belongs to a family—or if a described family should be recognized at all—are proposed and determined by practicing taxonomists. There are no hard rules for describing or recognizing a family, but in plants, they can be characterized on the basis of both vegetative and reproductive features of plant species. Taxonomists often take different positions about descriptions, and there may be no broad consensus across the scientific community for some time. The publishing of new data and opini ...
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Allokotosauria
Allokotosauria is a clade of early archosauromorph reptiles from the Middle to Late Triassic known from Asia, Africa, North America and Europe. Allokotosauria was first described and named when a new monophyletic grouping of specialized herbivorous archosauromorphs was recovered by Sterling J. Nesbitt, John J. Flynn, Adam C. Pritchard, J. Michael Parrish, Lovasoa Ranivoharimanana and André R. Wyss in 2015. The name Allokotosauria is derived from Greek meaning "strange reptiles" in reference to unexpected grouping of early archosauromorph with a high disparity of features typically associated with herbivory. History Nesbitt ''et al.'' (2015) defined the group as a stem-based taxon containing ''Azendohsaurus madagaskarensis'' and '' Trilophosaurus buettneri'' and all taxa more closely related to them than to ''Tanystropheus longobardicus'', ''Proterosuchus fergusi'', ''Protorosaurus speneri'' or ''Rhynchosaurus articeps''. Therefore, Allokotosauria includes the families Azendoh ...
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Sauropoda
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their body), and four thick, pillar-like legs. They are notable for the enormous sizes attained by some species, and the group includes the largest animals to have ever lived on land. Well-known genera include ''Brachiosaurus'', ''Diplodocus'', ''Apatosaurus'' and ''Brontosaurus''. The oldest known unequivocal sauropod dinosaurs are known from the Early Jurassic. ''Isanosaurus'' and ''Antetonitrus'' were originally described as Triassic sauropods, but their age, and in the case of ''Antetonitrus'' also its sauropod status, were subsequently questioned. Sauropod-like sauropodomorph tracks from the Fleming Fjord Formation (Greenland) might, however, indicate the occurrence of the group in the Late Triassic. By the Late Jurassic (150 million yea ...
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Convergent Evolution
Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last common ancestor of those groups. The cladistic term for the same phenomenon is homoplasy. The recurrent evolution of flight is a classic example, as flying insects, birds, pterosaurs, and bats have independently evolved the useful capacity of flight. Functionally similar features that have arisen through convergent evolution are ''analogous'', whereas '' homologous'' structures or traits have a common origin but can have dissimilar functions. Bird, bat, and pterosaur wings are analogous structures, but their forelimbs are homologous, sharing an ancestral state despite serving different functions. The opposite of convergence is divergent evolution, where related species evolve different traits. Convergent evolution is similar to parallel evo ...
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Plesiomorphy
In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, and synapomorphy, all mean a trait shared between species because they share an ancestral species. Apomorphic and synapomorphic characteristics convey much information about evolutionary clades and can be used to define taxa. However, plesiomorphic and symplesiomorphic characteristics cannot. The term ''symplesiomorphy'' was introduced in 1950 by German entomologist Willi Hennig. Examples A backbone is a plesiomorphic trait shared by birds and mammals, and does not help in placing an animal in one or the other of these two clades. Birds and mammals share this trait because both clades are descended from the same far distant ancestor. Other clades, e.g. snakes, lizards, turtles, fish, frogs, all have backbones and none are either birds n ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Synapomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis T ...
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Sauropodomorpha
Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had long necks and tails, were quadrupedal, and became the largest animals to ever walk the Earth. The '' prosauropods,'' which preceded the sauropods, were smaller and were often able to walk on two legs. The sauropodomorphs were the dominant terrestrial herbivores throughout much of the Mesozoic Era, from their origins in the Late Triassic (approximately 230 Ma) until their decline and extinction at the end of the Cretaceous. Description Sauropodomorphs were adapted to browsing higher than any other contemporary herbivore, giving them access to high tree foliage. This feeding strategy is supported by many of their defining characteristics, such as: a light, tiny skull on the end of a long neck (with ten or more elongated cervical vertebrae) ...
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