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Aspicilia Pacifica
''Aspicilia pacifica'' (pacific sunken disk lichen) is a white to grayish, brownish, or ocher crustose areolate lichen that commonly grows on siliceous rock or basalt along the seashore and in higher coastal mountains of California and Baja California.Field Guide to California Lichens, Stephen Sharnoff, Yale University Press, 2014, Aspicilia pacifica, Lichen Flora of the Greater Sonoran Desert Region. Vol 3, Nash, T.H., Ryan, B.D., Gries, C., Bugartz, F., (eds.) 2001/ref> It has numerous small (0.1–.8 mm), round to angular apothecia toward the middle of the thallus, with concave to flat black discs that are sometimes lightened with white pruina. Lichen spot test on the cortex and medulla are I−, K+ yellow to red, P+ orange, and C−. Secondary metabolites include much stictic acid, and some norstictic acid Norstictic acid is a depsidone produced as a secondary metabolites in lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria livin ...
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Crustose Lichen
Crustose lichens are lichens that form a crust which strongly adheres to the Substrate (biology), substrate (soil, rock, tree bark, etc.), making separation from the substrate impossible without destruction. The basic structure of crustose lichens consists of a cortex (botany), cortex layer, an algal layer, and a medulla. The upper cortex layer is differentiated and is usually pigmented. The algal layer lies beneath the cortex. The medulla fastens the lichen to the substrate and is made up of Fungus, fungal hyphae. The surface of crustose lichens is characterized by branching cracks that periodically close in response to climatic variations such as alternate wetting and drying regimes. Subtypes * Powdery – considered as the simplest subtype due to the absence of an organized thallus. :The thallus appears powdery. :E.g. Genera ''Lepraria'', ''Vezdaea'' * Endolithic – grows inside the rock, usually in interstitial spaces between mineral grains. The :upper cortex is usually d ...
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Lichen Spot Test
A spot test in lichenology is a spot analysis used to help identify lichens. It is performed by placing a drop of a chemical on different parts of the lichen and noting the colour change (or lack thereof) associated with application of the chemical. The tests are routinely encountered in dichotomous keys for lichen species, and they take advantage of the wide array of lichen products produced by lichens and their uniqueness among taxa. As such, spot tests reveal the presence or absence of chemicals in various parts of a lichen. They were first proposed by the botanist William Nylander in 1866. Three common spot tests use either 10% aqueous KOH solution (K test), saturated aqueous solution of bleaching powder or calcium hypochlorite (C test), or 5% alcoholic ''p''-phenylenediamine solution (P test). The colour changes occur due to presence of particular secondary metabolites in the lichen. There are several other less frequently used spot tests of more limited use that are employed ...
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Lichens Of North America
Irwin M. Brodo (born 1935) is an emeritus scientist at the Canadian Museum of Nature, in Ottawa, Ontario, Canada. He is an authority on the identification and biology of lichens. Irwin Brodo was honored in 1994 with an Acharius Medal presented to him by the International Association for Lichenology. Brodo did his undergraduate studies at Columbia University, and received a master's degree from Cornell University. He earned a Ph.D. in lichenology under the supervision of Henry Imshaug at Michigan State University. He later went on to teach at Université Laval and the University of Alaska, and he also supervised master's students at the University of Ottawa and Carleton University. Brodo's list of publications includes 75 research papers, 8 popular articles, 22 reviews and 6 editorials and obituaries. In 1993, Brodie was awarded the Mary E. Elliot Service Award for his meritorious service to the Canadian Botanical Association. One of Irwin Brodo's great achievements was the public ...
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Lichen Species
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

Norstictic Acid
Norstictic acid is a depsidone produced as a secondary metabolites in lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Lactones Phenols Lichen products Heterocyclic compounds with 4 rings {{lichen-stub ...
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Stictic Acid
Stictic acid is an aromatic organic compound, a product of secondary metabolism in some species of lichens. Stictic acid is the subject of preliminary biomedical research. Stictic acid has cytotoxic and apoptotic effects ''in vitro''. Computational studies suggest stictic acid may also stimulate p53 p53, also known as Tumor protein P53, cellular tumor antigen p53 (UniProt name), or transformation-related protein 53 (TRP53) is a regulatory protein that is often mutated in human cancers. The p53 proteins (originally thought to be, and often s ... reactivation. References Phenolic acids Phenol ethers Oxygen heterocycles Lactones Lichen products {{aromatic-compound-stub ...
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Secondary Metabolite
Secondary metabolites, also called specialised metabolites, toxins, secondary products, or natural products, are organic compounds produced by any lifeform, e.g. bacteria, fungi, animals, or plants, which are not directly involved in the normal growth, development, or reproduction of the organism. Instead, they generally mediate ecological interactions, which may produce a selective advantage for the organism by increasing its survivability or fecundity. Specific secondary metabolites are often restricted to a narrow set of species within a phylogenetic group. Secondary metabolites often play an important role in plant defense against herbivory and other interspecies defenses. Humans use secondary metabolites as medicines, flavourings, pigments, and recreational drugs. The term secondary metabolite was first coined by Albrecht Kossel, a 1910 Nobel Prize laureate for medicine and physiology in 1910. 30 years later a Polish botanist Friedrich Czapek described secondary metabolit ...
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Medulla (lichenology)
The medulla is a horizontal layer within a lichen thallus. It is a loosely arranged layer of interlaced hyphae below the upper cortex and photobiont A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Galloway, D.J. (1992). Flora of Australia - ''Lichen Glossary'' The medulla generally has a cottony appearance. It is the widest layer of a heteromerous lichen thallus.


References

Fungal morphology and anatomy Lichenology {{lichen-stub ...
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Cortex (lichen)
In botany, a cortex is an outer layer of a stem or root in a vascular plant, lying below the epidermis but outside of the vascular bundles. The cortex is composed mostly of large thin-walled parenchyma cells of the ground tissue system and shows little to no structural differentiation. The outer cortical cells often acquire irregularly thickened cell walls, and are called collenchyma cells. Plants Stems and branches In the three dimensional structure of herbaceous stems, the epidermis, cortex and vascular cambium form concentric cylinders around the inner cylindrical core of pith. Some of the outer cortical cells may contain chloroplasts, giving them a green color. They can therefore produce simple carbohydrates through photosynthesis. In woody plants, the cortex is located between the periderm (bark) and the vascular tissue (phloem, in particular). It is responsible for the transportation of materials into the central cylinder of the root through diffusion and may also be ...
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Pruina
Pruinescence , or pruinosity, is a "frosted" or dusty-looking coating on top of a surface. It may also be called a pruina (plural: ''pruinae''), from the Latin word for hoarfrost. The adjectival form is pruinose . Entomology In insects, a "bloom" caused by wax particles on top of an insect's cuticle covers up the underlying coloration, giving a dusty or frosted appearance. The pruinescence is commonly white to pale blue in color but can be gray, pink, purple, or red; these colors may be produced by Tyndall scattering of light. When pale in color, pruinescence often strongly reflects ultraviolet. Pruinescence is found in many species of Odonata, particularly damselflies of the families Lestidae and Coenagrionidae, where it occurs on the wings and body. Among true dragonflies it is most common on male Libellulidae (skimmers). In the common whitetail and blue dasher dragonflies (''Plathemis lydia'' and ''Pachydiplax longipennis''), males display the pruinescence on the back of th ...
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Areolate
Lichens are composite organisms made up of multiple species: a fungal partner, one or more photosynthetic partners, and sometimes a basidiomycete yeast. They are regularly grouped by their external appearance – a characteristic known as their growth form. Lichenologists have described a dozen of these forms: areolate, byssoid, calicioid, cladoniform, crustose, filamentous, foliose, fruticose, gelatinous, leprose, placoidioid and squamulose. Of these, crustose, foliose and fruticose are the most commonly encountered. With the exception of calicioid lichens, growth forms are based on the appearance of the thallus, which is the vegetative (non-reproductive) part of the lichen. In most species, this form is determined by the lichen's fungal partner, though in a small number, it is instead the photobiont that determines the lichen's morphology. In some growth forms, the outermost layer of the thallus consists of tightly woven fungal . This layer, known as the cortex, may be found ...
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Thallus
Thallus (plural: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. Many of these organisms were previously known as the thallophytes, a polyphyletic group of distantly related organisms. An organism or structure resembling a thallus is called thalloid, thallodal, thalliform, thalline, or thallose. A thallus usually names the entire body of a multicellular non-moving organism in which there is no organization of the tissues into organs. Even though thalli do not have organized and distinct parts (leaves, roots, and stems) as do the vascular plants, they may have analogous structures that resemble their vascular "equivalents". The analogous structures have similar function or macroscopic structure, but different microscopic structure; for example, no thallus has vascular tissue. In exceptional cases such as the Lemnoideae, where ...
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