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Arechia
''Arechia'' is an extinct genus of stingaree from the Eocene epoch. It is the oldest known stingaree genus, though molecular data suggests they diverged from Butterfly rays about 75 million years ago or from Deepwater stingrays around the time of the K-Pg transition. The type species, ''A. arambourgi'', is known from isolated teeth found in the Ypresian deposits of Ouled Abdoun, Morocco. It is named for Camille Arambourg, who originally ascribed these teeth to ''Raja praealba'' in 1952. Henri Cappetta revised the species in 1983, erecting this genus for those Arambourg called males in this genus and placed the ones he called females in '' Merabatis''. The second species, ''A. crassicaudata'' is from the Ypresian-age Bolca Lagerstätte of Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymo ...
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Animalia
Animals are multicellular, eukaryotic organisms in the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of cells, the blastula, during embryonic development. Over 1.5 million living animal species have been described—of which around 1 million are insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a bilaterally symmetric body plan. The Bilateria include the protostomes, containing animals such as nematodes, arthropods, flatworms, annelids and molluscs, and the deuterostomes, containing the echinode ...
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Ypresian
In the geologic timescale the Ypresian is the oldest age (geology), age or lowest stage (stratigraphy), stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by the Eocene Lutetian Age. The Ypresian is consistent with the lower Eocene. Events The Ypresian Age begins during the throes of the Paleocene–Eocene Thermal Maximum (PETM). The Fur Formation in Denmark, the Messel shales in Germany, the Oise amber of France and Cambay amber of India are of this age. The Eocene Okanagan Highlands are an uplands subtropical to temperate series of lakes from the Ypresian. Stratigraphic definition The Ypresian Stage was introduced in scientific literature by Belgium, Belgian geologist André Hubert Dumont in 1850. The Ypresian is named after the Flanders, Flemish city of Ypres in Belgium (spelled ''Ieper'' in Dutch). The definitions of the original stage were totally different from the modern ones. The Ypresi ...
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Monte Bolca
Monte Bolca is a lagerstätte near Verona, Italy that was one of the first fossil sites with high quality preservation known to Europeans, and is still an important source of fossils from the Eocene. Geology Monte Bolca was uplifted from the Tethys Ocean floor during the formation of the Alps, in two stages: one 24 million years ago, and one between 30 and 50 million years ago. The entire formation consists of of limestone, all of which contain fossils, but interspersed in which are the lagerstätte layers that contain the highly preserved specimens. Within these layers, the fish and other specimens are so highly preserved that their organs are often completely intact in fossil form, and even the skin colorWilliams, MattTaphonomy of Monte Bolca University of Bristol can sometimes be determined. The normal rearrangement of the specimens caused by mud-dwelling organisms in the layer before it turned to stone has been avoided—it is assumed that the mud in question was low in ox ...
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Henri Cappetta
Henri Cappetta is a French ichthyologist specializing in the paleontology of shark Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachimo ...s and rays. References * H. Cappetta, Handbook of Paleoichthyology (Gustav Fischer, 1987) External links Henri Cappetta on www.isem.cnrs.fr(French) New sharks and rays from the Cenomanian and Turonian of Charentes, France. Romain Vullo, Henri Cappetta and Didier Néraudeau, Acta Palaeontol. Pol. 52 (1), pp. 99–116, 2007 French ichthyologists Living people Year of birth missing (living people) {{France-zoologist-stub ...
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Raja (fish)
''Raja'', also known as ''raia'', is a genus of Skate (fish), skates in the family (biology), family Rajidae containing 16 species. Formerly a wastebasket genus, many species historically placed here have been moved to other genera in the family, such as ''Amblyraja'', ''Beringraja'', ''Dipturus'', ''Leucoraja'' and ''Rostroraja''. ''Raja'' are flat-bodied, cartilaginous fish with a rhombic shapes due to their large pectoral fins extending from or nearly from the snouts to the bases of their tails. Their sharp snouts are produced by a cranial projection of rostral cartilage. The mouth and gills are located on underside of the body. They may be either solid-coloured or patterned, and most skates have spiny or thorn-like structures on the upper surface, and some species contain weak electrical organs within their tails. Mating typically occurs in the spring and the female lays numerous Egg (biology), eggs per clutch which are encapsulated in leathery cases, commonly known as "Egg cas ...
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Camille Arambourg
Camille Arambourg ( February 3, 1885– November 19, 1969) was a French vertebrate paleontologist. He conducted extensive field work in North Africa. In the 1950s he argued against the prevailing model of Neanderthals as brutish and simian. During World War I he was in Military service. After that he was a professor of Geology at the Institut Agricole d'Alger, and after that a professor of Paleontology at Muséum National d'Histoire Naturelle in Paris, where he succeeded his teacher Marcellin Boule. The pterosaur ''Arambourgiania'' is named after him. He was President of the PanAfrican Archaeological Association from 1959 to 1963. Publications * (1942) "L’ Elephas recki ''Palaeoloxodon recki'' is an extinct species of elephant native to Africa during the Pliocene and Pleistocene. At up to 14 feet (4.27 metres) in shoulder height, it was one of the largest elephant species to have ever lived. It is believed that ... Dietrich. Exposition systématique et ses affinités". ...
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Ouled Abdoun Basin
The Oulad Abdoun Basin (also known as the Ouled Abdoun Basin or Khouribga Basin) is a phosphate sedimentary basin located in Morocco, near the city of Khouribga. It is the largest in Morocco, comprising 44% of Morocco's phosphate reserves, and at least 26.8 billion tons of phosphate. It is also known as an important site for vertebrate fossils, with deposits ranging from the Late Cretaceous (Cenomanian-Turonian) to the Eocene epoch (Ypresian), a period of about 25 million years. Geography The Oulad Abdoun is located west of the Atlas Mountains, near the city of Khouribga. The Oulad Abdoun phosphate deposits encompass some , an area of . The Oulad Abdoun is the largest and northernmost of Morocco's major phosphate basins, which from northeast to southwest, include the Ganntour, Meskala, and Oued Eddahab (Laayoune-Baa) basins. Paleobiota The Oulad Abdoun Basin stretches from late Cretaceous to the Eocene, and contains abundant marine vertebrate fossils, including sharks, bo ...
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Cretaceous–Paleogene Extinction Event
The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million years ago. With the exception of some ectothermic species such as sea turtles and crocodilians, no tetrapods weighing more than survived. It marked the end of the Cretaceous Period, and with it the Mesozoic era, while heralding the beginning of the Cenozoic era, which continues to this day. In the geologic record, the K–Pg event is marked by a thin layer of sediment called the K–Pg boundary, which can be found throughout the world in marine and terrestrial rocks. The boundary clay shows unusually high levels of the metal iridium, which is more common in asteroids than in the Earth's crust. As originally proposed in 1980 by a team of scientists led by Luis Alvarez and his son Walter, it is now generally thought that the K–Pg extinction was cause ...
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Chordata
A chordate () is an animal of the phylum Chordata (). All chordates possess, at some point during their larval or adult stages, five synapomorphies, or primary physical characteristics, that distinguish them from all the other taxa. These five synapomorphies include a notochord, dorsal hollow nerve cord, endostyle or thyroid, pharyngeal slits, and a post-anal tail. The name “chordate” comes from the first of these synapomorphies, the notochord, which plays a significant role in chordate structure and movement. Chordates are also Bilateral symmetry, bilaterally symmetric, have a coelom, possess a circulatory system, and exhibit Metameric, metameric segmentation. In addition to the morphological characteristics used to define chordates, analysis of genome sequences has identified two conserved signature indels (CSIs) in their proteins: cyclophilin-like protein and mitochondrial inner membrane protease ATP23, which are exclusively shared by all vertebrates, tunicates and cep ...
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Plesiobatid
The deepwater stingray or giant stingaree (''Plesiobatis daviesi'') is a species of stingray and the sole member of the family Plesiobatidae. It is widely distributed in the Indo-Pacific, typically over fine sediments on the upper continental slope at depths of . This species reaches in length and in width. It has an oval pectoral fin disc with a long, flexible, broad-angled snout. Most of the entire latter half of its tail supports a distinctively long, slender, leaf-shaped caudal fin. Its coloration is dark above and white below, and its skin is almost completely covered by tiny dermal denticles. Preying on crustaceans, cephalopods, and bony fishes, the deepwater stingray may hunt both on the sea floor and well above it in open water. It is probably aplacental viviparous, with the mother supplying her gestating young with histotroph ("uterine milk"). Captured rays merit caution due to their long, venomous stingers. This species is taken by deepwater commercial fisheries, b ...
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Gymnurid
The butterfly rays are the rays forming the genus ''Gymnura'' and the family Gymnuridae. They are found in warm oceans worldwide, and occasionally in estuaries. The body of butterfly rays is flattened and surrounded by an extremely broad disc formed by the pectoral fins, which merge in front of the head. They have a very short, thread-like, tail. They are up to in width. McEachran ''et al.'' place the butterfly rays in the subfamily Gymnurinae of the family Dasyatidae, but this article follows FishBase and ITIS in treating them as a family. Species There are currently 12 species in this genus (others are considered synonyms): * '' Gymnura altavela'' (Linnaeus, 1758) – Spiny butterfly ray * '' Gymnura australis'' ( E. P. Ramsay & Ogilby, 1886) – Australian butterfly ray * ''Gymnura crebripunctata'' ( W. K. H. Peters, 1869) – Longsnout butterfly ray * '' Gymnura japonica'' (Temminck & Schlegel, 1850) – Japanese butterfly ray * ''Gymnura marmorata'' ( J. G. Cooper, ...
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